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Agama rueppelli VAILLANT, 1882

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Higher TaxaAgamidae (Agaminae), Sauria, Iguania, Squamata (lizards) 
SubspeciesAgama rueppelli rueppelli VAILLANT in REVOIL 1882
Agama rueppelli occidentalis PARKER 1932
Agama rueppelli septentrionalis PARKER 1932 
Common NamesRüppell’s Agama 
SynonymAgama ruppelli VAILLANT in REVOIL 1882: 6 (in error)
Agama vaillanti BOULENGER 1895: 12 (fide LOVERIDGE 1932)
Agama rueppelli — BOULENGER 1885: 355 (nom. emend.)
Agama Rueppellii — BOULENGER 1897: 130
Agama vaillanti — LÖNNBERG 1911
Agama rueppelli — WERNER 1913: 9
Agama vaillanti — LOVERIDGE 1920: 142
Agama vaillanti — SCORTECCI 1931: 141
Agama rueppelli — PARKER 1932: 352
Agama rueppelli — WERMUTH 1967: 22
Agama rueppelli — LANZA 1990
Agama rueppelli — WAGNER & BAUER 2011
Agama ruppelli — SPAWLS et al. 2018: 235

Agama rueppelli occidentalis PARKER 1932
Agama rueppelli occidentalis — PARKER 1936: 604
Agama rueppelli occidentalis — DREWES 1972

Agama rueppelli septentrionalis PARKER 1932
Agama ruppelli septentrionalis — LOVERIDGE 1957: 193
Agama rueppelli septentrionalis — WERMUTH 1967: 23 
DistributionSomalia, E Ethiopia, Kenya, S Sudan

occidentalis: “South-Abessinia [= Ethiopia] through N Kenya”. Type locality: mouth of the Kaliokwell River, Lake Rudolf, Kenya Colony.

septentrionalis: E Kenya. Type locality: Mount Njero (Njiro), Madago’s Village, Voi and Mbunji, Kenya Colony. Paratypes: MCZ

vaillanti: Somalia; Type locality: “Ogaden and between Laffarug and Aberio, Somaliland”.

Type locality: Bender-Meraya, Pays Comalis [= Somalia]  
Reproductionoviparous 
TypesSyntypes. MNHN 5897; BMNH 1946.8.24.50
Lectotype [vaillanti]: MSNG 28850, adult male, from ‘‘Ogaden,’’ Ethiopia.
Syntypes: MCZ, USNM, CAS [septentrionalis]
Syntypes: BMNH, USNM [occidentalis] 
DiagnosisDiagnosis. A medium-sized Agama with a maximum total length of 280 mm. Males (58–88 mm SVL) smaller than females (76– 88 mm SVL). Body depressed, hind limbs long. Tail more than 65% of total length, very broad at its base, tapering sharply. Gular pouch absent. Head convex, with the upper surface of the snout flat. Head scales smooth, equal to ventral scales, with the occipital scales variable in size (equal to other head scales to slightly enlarged). Nostril directed posterodorsally, completely visible from above, pierced in the posterior part of a large nasal scale, situated above the canthus rostralis. Usually with one to two scales between nasal and first supraciliary scales. Supraciliary scales smooth. Nuchal and dorsal crest absent. Ear opening only half the size of the eye, surrounded by tufts of spinose scales hiding the tympanum. Body scales large, homogeneous, keeled, mucro- nate, and erect, in 54–64 scale rows around midbody. Vertebral scales 59–66. Gular and ventral scales smooth, ventrals smaller than dorsal scales. Lamellae 14–16 under the fourth toe, third and fifth toes nearly equal in length. Males with one row of 9–13 precloacal pores. 
Comment 
EtymologyNamed after Eduard Rüppell (1794-1884), German herpetologist. 
References
  • Boettger,O. 1893. Übersicht der von Prof. C. Keller anlässlich der Ruspoli'schen Expedition nach den Somaliländern gesammelten Reptilien und Batrachier. Zool. Anz. 16 (416): 113-119 - get paper here
  • Böhme, W. 1985. Zoogeographical patterns of the lizard fauna of the African subsaharan savanna belt, with preliminary description of a new chameleon. In: Schuchmann, Karl-L. (Hrsg.): Proceedings of the International Symposium on African Vertebrates. Systematics, Phylogeny and Evolutionary Ecology. - Museum A. Koenig (Bonn), Vol. 4: 471-478.
  • Boulenger, G. A. 1898. On a second collection of reptiles made by Mr. E. Lord-Phillips in Somaliland. Ann. Mag. nat. Hist. (7) 2: 130-133 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the Lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger,G.A. 1895. Rettili e Batraci. in, Esplorazione del Giuba e dei suoi Affluenti compiuta dal Cap. V. Bottego durante gli Anni 1892-93 sotto gli auspicii della Società Geografica Italiana. Annali Mus. civ. Stor. nat. Giacomo Doria (2) 15: 9-18
  • Cochran, Doris M. 1961. Type specimens of reptiles and amphibians in the United States National Museum. Bull. US Natl. Mus. (220): xvi + 291 pp. - get paper here
  • Drewes, R.C. 1972. Report on a collection of reptiles and amphibians from the Ilemi Triangle, Southwestern Sudan. Occ. Pap. Cal. Acad. Sci. (100): 1-14 - get paper here
  • Lanza, B. 1990. Amphibians and reptiles of the Somali Democratic Republic: check list and biogeography. Biogeographia, 14: 407-465 [1988]
  • Largen, M.J.; Spawls, S. 2006. Lizards of Ethiopia (Reptilia Sauria): an annotated checklist, bibliography, gazetteer and identification. Tropical Zoology 19 (1): 21-109 - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • LEACHÉ, Adam D.; Rebecca A. CHONG, Theodore J. PAPENFUSS, Philipp WAGNER, Wolfgang BÖHME, Andreas SCHMITZ, Mark-Oliver RÖDEL, Matthew LEBRETON, Ivan INEICH, Laurent CHIRIO, Aaron BAUER, Edem A. ENIANG, & Sherif BAHA EL DIN 2009. Phylogeny of the genus Agama based on mitochondrial DNA sequence data. Bonner zoologische Beiträge 56 (4): 273–278 - get paper here
  • Lönnberg, E. 1911. Reptiles, Batrachians and Fishes collected by the swedish Zoological Expedition to British East Africa 1911. Kungl. Sv. Vet. Akademiens Handligar. 47(6): 1-24.
  • Loveridge, A. 1957. Check list of the reptiles and amphibians of east Africa (Uganda, Kenya, Tanganyika, Zanzibar). Bull. Mus. Comp. Zool. Harvard 117 (2): 153-362 - get paper here
  • Loveridge,A. 1920. Notes on East African lizards collected 1915-1919, with description of a new genus and species of skink and new subspecies of gecko. Proc. Zool. Soc. London 1920: 131-167 - get paper here
  • Mazuch, Tomáš 2013. Amphibians and Reptiles of Somaliland and Eastern Ethiopia. Tomáš Mazuch Publishing, 80 pp. ISBN: 978-80-905439-0-4 - get paper here
  • Parker, H. W. 1942. The lizards of British Somaliland. Bull. Mus. Comp. Zool. Harvard 91: 1—101 - get paper here
  • Parker,H.W. 1932. Scientific results of the Cambridge expedition to the east African lakes, 1930-31. 5. Reptiles and amphibians. Zool. J. Linnean Soc. 38: 213-229
  • Parker,H.W. 1932. Two collections af amphibians and reptiles from British Somaliland. Proc. Zool. Soc. London 1932: 335-367
  • Parker,H.W. 1936. Reptiles and amphibians collected by the Lake Rudolf Rift Valley Expedition, 1934. Ann. Mag. nat. Hist. (10) 18: 594-609
  • Scortecci, G. 1930. Secondo contributo alla conoscenza dei rettili della Somalia Italiana. Atti della Società Italiana di Scienze Naturali e del Museo Civico di Storia Naturale in Milano, 69:127—152.
  • Spawls, S. & Rotich, D. 1997. An annotated checklist of the lizards of Kenya. J. East African Nat. Hist. 86: 61-83 - get paper here
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Vaillant, M.L. 1882. Reptiles et Batraciens. in: Révoil: Mission G. Révoil aux pays Comalis, Faune et Flore. Faune et Flore des Pays Comalis, Afr. orient., 25 pp, 3 plates
  • Wagner, P. 2014. A new cryptic species of the Agama lionotus complex from south of the Ngong Hills in Kenya. Salamandra 50 (4): 187-200 - get paper here
  • Wagner, Philipp and Aaron Bauer 2011. A new dwarf Agama (Sauria: Agamidae) from Ethiopia. Breviora (527): 1-19 - get paper here
  • Werner, F. 1913. Neue oder seltene Reptilien und Frösche des Naturhistorischen Museums in Hamburg. Reptilien der Ostafrika-Expedition der Hamburger Geographischen Gesellschaft 1911/12. Leiter: Dr. E. Obst. Reptilien und Amphibien von Formosa. Jb. Hamb. wiss. Anst., 30 [1912], 2. Beiheft: 1-39, 40-45, 45-51 - get paper here
 
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