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Anolis campbelli (KÖHLER & SMITH, 2008)

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common Names 
SynonymNorops campbelli KÖHLER & SMITH 2008
Norops campbelli — NICHOLSON et al. 2012 
DistributionW Guatemala (Sierra de los Cuchumatanes, western Guatemala, from 1540 to 1660 m elevation).

Type locality: trail to Laguna Yolnabaj, Aldea Yalambojoch, Municipalidad Nentón, Departamento de Huehuetenango, Guatemala, 1540 m elevation, 16° 00’ 17’’ N, 91° 34’ 05’’ W. Map legend:
Type locality - Type locality.
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: UTA R-46038, an adult male, University of Texas at Arlington, collected 29 May 1998 by Manuel Acevedo and Marco V. Centeno. Original number MEA 1436. 
CommentDiagnosis.—A medium-sized species (snout– vent length [SVL] in largest specimen 51.3 mm) of the genus Norops (sensu Guyer and Savage, 1987, 1992) that belongs to the Norops schiedei species group (sensu NietoMontes de Oca, 2001). Norops campbelli differs from species of the N. schiedei group by the following characteristics (condition for N. campbelli in parentheses): Norops cobanensis: hemipenis small and unilobate (large and bilobate); scales comprising supraorbital semicircles usually distinctly larger than scales between semicircles (about subequal); smooth or only some faintly keeled midventral scales (most mid-ventral scales weakly, yet definitely keeled); some keels usually oblique or oriented transversely immediately anterior to prefrontal depression (oriented nearly longitudinally); scales in prefrontal depression mostly smooth (all scales in prefrontal depression strongly keeled); circumorbital row usually complete (rudimentary or almost absent). Norops cuprinus: prenasal distinct (prenasal fused with circumnasal); lorilabial row usually complete (usually incomplete). Norops cymbobs: para-vertebral scales slightly to moderately, albeit distinctly larger than scales in adjacent rows (paravertebral scales not larger than scales in adjacent rows); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); few or most preantebrachials and supra-antebrachials multicarinate (all unicarinate); prenasal distinct (prenasal fused with circumnasal). Norops duellmani: prenasal distinct (prenasal fused with circumnasal). Norops hobartsmithi: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal); scales in prefrontal depression mostly smooth (all scales in prefrontal depression strongly keeled); circumorbital row usually complete (rudimentary or almost absent); lorilabial row usually complete (usually incomplete). Norops johnmeyeri: postanal scales moderately enlarged in males (not enlarged); dewlap well developed in both sexes (not developed in females); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops matudai: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal); lorilabial row usually complete (usually incomplete). Norops milleri: paravertebral scales slightly to moderately, albeit distinctly, larger than scales in adjacent rows (paravertebral scales not larger than scales in adjacent rows); smooth or only some faintly keeled midventral scales (most mid-ventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops naufragus: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); middorsal scales between levels of axilla and groin 82–100 (68–69); few or most preantebrachials and supra-antebrachials multicarinate (all unicarinate). Norops parvicirculatus: scales comprising supraorbital semicir cles distinctly smaller than scales between semicircles (about subequal); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal); circumorbital row usually complete (rudimentary or almost absent). Norops pijolensis: middorsal scales between levels of axilla and groin 43–54 (68–69); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops polyrhachis: smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled). Norops purpurgularis: horizontal loreal rows 7–9 (5–6); middorsal scales between levels of axilla and groin 52–62 (68– 69); smooth or only some faintly keeled midventral scales (most midventral scales weakly, yet definitely keeled); prenasal distinct (prenasal fused with circumnasal). Norops schiedei: some keels usually oblique or oriented transversely immediately anterior to prefrontal depression and in depression (all head keels oriented nearly longitudinally).

Species group: Norops auratus Species Group (fide Nicholson et al. 2012) 
EtymologyEtymology.—The specific epithet honors the foremost student of Guatemalan herpetology, Jonathan Campbell, American herpetologist, who also collected one of the type specimens and photographed the only two known specimens. 
References
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Köhler, Gunther and Eric N. Smith 2008. A New Species of Anole of the Norops schiedei group from Western Guatemala (Squamata: Polychrotidae). Herpetologica 64 (2): 216-223 - get paper here
  • Murphy, J.B. 1997. A lok at the life of Jonathan A. Campbell. Fauna 1 (1): 72-78
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
 
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