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Anolis frenatus COPE, 1899

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common NamesBridled Anole 
SynonymAnolis frenatus COPE 1899: 6
Anolis frenatus — DUNN 1937: 9
Anolis purpurescens — TAYLOR 1956
Anolis frenatus — PETERS & OREJAS-MIRANDA 1970: 53
Dactyloa frenata — GUYER & SAVAGE 1986
Dactyloa frenata — KÖHLER 2000: 59
Dactyloa frenata — NICHOLSON et al. 2012 
DistributionCosta Rica, Panama, Colombia (Caribbean), elevation up to 900 m.

Type locality: unknown; Barbour (1934) suggested Baranquilla, Colombia; exact location uncertain according to Savage and Talbot (1978).  
Reproductionoviparous 
TypesHolotype: Lost (fide LOTZKAT et al. 2013) 
CommentDistribution: Reports from Venezuela are either in error or based on misidentifications (UGUETO et al. 2009).

Savage and Talbot (1978) reported that one of the two specimens of Peters’ original description of A. squamulatus was in fact an A. frenatus mistakenly believed to have come from Puerto Cabello, Venezuela.

Genomics: Marc Tollis et al. (Arizona State University) presented a draft genome assembly of this species at SICB 2014 (see link).

Species group: Dactyloa latifrons species group (NICHOLSON et al. 2012). 
References
  • Barbour, T. 1934. The anoles. II. The mainland species from México southward. Bull. Mus. Comp. Zool. Harvard 77: 121-155. - get paper here
  • BATISTA, ABEL; MILAN VESELY, KONRAD MEBERT, SEBASTIAN LOTZKAT & GUNTHER KÖHLER 2015. A new species of Dactyloa from eastern Panama, with comments on other Dactyloa species present in the region. Zootaxa 4039 (1): 057–084 - get paper here
  • Cope, E.D. 1899. Contributions to the herpetology of New Granada and Argentina, with descriptions of new forms. Philadelphia Mus. Sci. Bull. (1): 1-19
  • Crandell, Kristen E.; Anthony Herrel, Mahmood Sasa, Jonathan B. Losos, Kellar Autumn 2014. Stick or grip? Co-evolution of adhesive toepads and claws in Anolis lizards. Zoology 117 (6): 363-369 - get paper here
  • Dunn, Emmett R. 1937. The giant mainland anoles. Proc. New England zool. Club 16: 5-9
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • KÖHLER, GUNTHER 2014. Characters of external morphology used in Anolis taxonomy—Definition of terms, advice on usage, and illustrated examples. Zootaxa 3774 (2): 201–257 - get paper here
  • LOTZKAT, SEBASTIAN; ANDREAS HERTZ, JOE-FELIX BIENENTREU & GUNTHER KÖHLER 2013. Distribution and variation of the giant alpha anoles (Squamata: Dactyloidae) of the genus Dactyloa in the highlands of western Panama, with the description of a new species formerly referred to as D. microtus. Zootaxa 3626 (1): 1–54 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • Savage, J.M. 2002. The Amphibians and Reptiles of Costa Rica: A Herpetofauna Between Two Continents, Between Two Seas. University of Chicago Press, 934 pp. [review in Copeia 2003 (1): 205]
  • Savage, Jay M. & Talbot, James J. 1978. The giant Anoline lizards of Costa Rica and Western Panama. Copeia 1978 (3): 480-492 - get paper here
  • Tollis, Marc; Elizabeth D. Hutchins, Jessica Stapley, Shawn M. Rupp, Walter L. Eckalbar, Inbar Maayan, Eris Lasku, Carlos R. Infante, Stuart R. Dennis, Joel A. Robertson, Catherine M. May, Michael R. Crusoe, Eldredge Bermingham, Dale F. DeNardo, S. T 2018. Comparative Genomics Reveals Accelerated Evolution in Conserved Pathways during the Diversification of Anole Lizards https://academic.oup.com/gbe/advance-article-abstract/doi/10.1093/gbe/evy013/4817506 - get paper here
  • Ugueto, Gabriel N.; Gilson Rivas, Tito Barros and Eric N. Smith 2009. A Revision of the Venezuelan Anoles II: Redescription of Anolis squamulatus Peters 1863 and Anolis tigrinus Peters 1863 (Reptilia: Polychrotidae). Carib. J. Sci. 45 (1): 30-51 - get paper here
 
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