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Anolis immaculogularis KÖHLER, TREJO PÉREZ, PETERSEN & MÉNDEZ DE LA CRUZ, 2014

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common Names 
SynonymAnolis immaculogularis KÖHLER, TREJO PÉREZ, PETERSEN & MÉNDEZ DE LA CRUZ 2014 
DistributionMexico (Oaxaca)

Type locality: Puerto Escondido, Punta Colorada (15.87038°N, 97.10152°W, WGS84), 40 m, Estado de Oaxaca, Mexico  
Reproductionoviparous 
TypesHolotype. SMF 96266, an adult male; collected 14 November 2012 by Gunther Köhler and Jesús García Grajales. Field tag number GK-4116.
Paratypes. All from Estado de Oaxaca, Mexico: SMF 96267, same collecting data as holotype. IBH 26567, from Puerto Escondido, Universidad del Mar (15.88934°N, 97.07703°W, WGS84), 80 m, collected 14 November 2012 by Gunther Köhler and Jesús García Grajales. IBH 26560–64, SMF 96263–64, from near Huatulco, 15.78191°N, 96.16257°W, 90 m; collected 12 November 2012 by Gunther Köhler and Raúl Gómez Trejo Pérez. IBH 26993–94, SMF 93842–43, 96467–68 from Parque National Huatulco, 15.73268°N, 96.16357°W, 10 m; collected 28 February 2013 by Gunther Köhler and Raúl Gómez Trejo Pérez. IBH 26562–63, SMF 93842–43, 96263–64, 96467 are males, IBH 26560, 26564, 26993, SMF 96267, 96468 are females, IBH 26561, 26567, 26569 are juveniles. 
CommentDiagnosis. A small to moderate-sized species (SVL in largest male 43.0 mm, largest female 49.5 mm) of the genus Anolis (sensu Poe 2004) that differs from all Mexican and Central American anoles except A. boulengerianus, A. carlliebi, A. quercorum, A. subocularis, and A. sacamecatensis by having a combination of (1) strongly keeled ventral scales; (2) usually a patch of three greatly enlarged supraocular scales; (3) 13–20 rows of slightly to moderately enlarged dorsal scales that are smaller than ventral scales; (4) moderately long hind legs, longest toe of adpressed hind leg usually reaching to level of posterior border of eye or beyond, ratio shank length/ SVL 0.24–0.31; and (5) a large pinkish red dewlap in males and a small red or pink dewlap in females. Anolis immaculogularis differs from A. carlliebi, A. quercorum, and A. sacamecatensis by having longer legs with the longest toe of adpressed hind leg usually reaching to level of posterior border of eye or beyond, ratio shank length/ SVL 0.24–0.31 (vs. to ear opening or to a point between ear opening and eye, ratio shank length/SVL 0.20–0.26). Anolis immaculogularis differs from A. subocularis in male dewlap coloration in life (pinkish red without paler areas around gorgetals in A. immaculogularis vs. pinkish red with paler areas around gorgetals in A. subocularis). Also, in A. immaculogularis, the subocular scales are usually broadly in contact with the supralabials (vs. subocular scales separated from supralabials by one scale row or these scales in contact in A. subocularis). Anolis immaculogularis differs from A. boulengerianus in male dewlap coloration in life (pinkish red without paler areas around gorgetals in A. immaculogularis vs. orange yellow with paler areas around gorgetals in A. boulengerianus). 
EtymologyThe name immaculogularis is a compound adjective derived from immaculosus (Latin for not spotted) and gula (Latin for throat) referring to male dewlap that lacks pale spots around the gorgetals, a diagnostic character to distinguish Anolis immaculogularis from its supposed closest relative, A. boulengerianus. 
References
  • KÖHLER, GUNTHER; RAÚL GÓMEZ TREJO PÉREZ, CLAUS BO P. PETERSEN & FAUSTO R. MÉNDEZ DE LA CRUZ 2014. A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species. Zootaxa 3862 (1): 001–210 - get paper here
  • KÖHLER, GUNTHER; RAÚL GÓMEZ TREJO PÉREZ, CLAUS BO P. PETERSEN & FAUSTO R. MÉNDEZ DE LA CRUZ 2015. A revision of the Mexican Anolis (Reptilia, Squamata, Dactyloidae) from the Pacific versant west of the Isthmus de Tehuantepec in the states of Oaxaca, Guerrero, and Puebla, with the description of six new species: Addendum. Zootaxa 3914 (4): 483–489 - get paper here
  • Kwet, Axel 2015. Liste der im Jahr 2014 neu beschriebenen Reptilien. Terraria-Elaphe 2015 (3): 50-64 - get paper here
  • Mata-Silva, Vicente, Jerry D. Johnson, Larry David Wilson and Elí García-Padilla. 2015. The herpetofauna of Oaxaca, Mexico: composition, physiographic distribution, and conservation status. Mesoamerican Herpetology 2 (1): 6–62 - get paper here
 
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