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Anolis meridionalis BOETTGER, 1885

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Higher TaxaDactyloidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common Names 
SynonymAnolis meridionalis BOETTGER 1885: 437
Anolis holotropis BOULENGER 1895: 522
Norops sladeniae BOULENGER 1903: 69
Anolis steinbachi GRIFFIN 1917 (fide VANZOLINI & WILLIAMS 1970)
Norops marmorata AMARAL 1932:63
Anolis chrysolepis meridionalis — HELLMICH 1960:22 (partim)
Anolis meridionalis — PETERS et al. 1970: 60
Anolis meridionalis — DIRKSEN & DE LA RIVA 1999
Norops meridionalis — NICHOLSON 2002
Anolis meridionalis — D’ANGIOLELLA et al. 2011
Norops meridionalis — NICHOLSON et al. 2012
Norops meridionalis — CACCIALI et al. 2016 
DistributionBrazil (Mato Grosso, Goias, São Paulo), Paraguay, Bolivia (Beni, Santa Cruz)

Type locality: Paraguay. Neotype locality: Colonia Ybycui, Estancia Ybycui, Departamento Canindeyu ́, Paraguay, (designation by Motte and Cacciali, 2009) Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesNeotype: MNHN, desgnated by MOTTE & CACCIALI 2009
Holotype: lost
For types of synonyms see D’ANGIOLELLA et al. 2011. 
CommentSynonymy after Peters et al. 1970, VANZOLINI & WILLIAMS 1970, and D’ANGIOLELLA et al. 2011.

N. meridionalis may be composed by at least five cryptic species (Guarnizo et al. 2005). However, they do not make any formal changes.

Description. Maximum SVL 56 mm. Specimens analyzed presented vertebral scales anteriorly small, keeled, posteriorly increasing in size and forming a gradual transition toward the granules on flanks; they do not form rows of enlarged scales along dorsum. The neotype description provided by Motte and Cacciali (2009) states that vertebral scales form eight rows of enlarged scales along back that are distinctly larger than scales on the flanks. Scales on upper arm keeled, markedly larger than vertebral scales. Scales on snout anteriorly weakly to distinctly keeled, uni- or multi- carinate and slightly larger than the flat posterior scales. Supraorbital semicircles with enlarged, weakly keeled scales. Suprao- cular region with a group of distinctly enlarged scales surrounded posteriorly and laterally by granules and anteriorly by smaller scales. Interparietal scale distinctly larger than adjacent scales. Loreals 6–7; canthals 5–6; 8–10 scales between second canthals; 0– 1 scales between supraorbital semicircles; 0– 2 scales between supraorbital semicircles and interparietal, postmentals 5–8 [from D’ANGIOLELLA et al. 2011].

Species group: Norops auratus Species Group (fide Nicholson et al. 2012) 
References
  • Araujo, Cybele O.; Fábio Maffei, Décio T. Corrêa, Guilherme M. Moya, Bruno T. M. Nascimento, Selma Maria A. Santos 2014. Lizards from Estação Ecológica de Santa Bárbara, a remnant of Cerrado in the state of São Paulo, Brazil. Check List 10 (5): 1038-10 - get paper here
  • Barbour, T. 1934. The anoles. II. The mainland species from México southward. Bull. Mus. Comp. Zool. Harvard 77: 121-155. - get paper here
  • Boettger,O. 1885. Berichtigung der Liste von Reptilien und Batrachiern aus Paraguay. Zeitschr. Naturw. 58: 436-437 - get paper here
  • Boulenger, George A. 1895. Description of a new Anolis from Brazil. Ann. Mag. nat. Hist. (6)15 (90): 522-523 - get paper here
  • Boulenger, George A. 1903. List of the batrachians and reptiles collected by M. A. Robert at Chapadá, Matto Grosso, and presented by Mrs. Percy Sladen to the British Museum. (Percy Sladen Expedition to Central Brazil). Proceedings of the Zoological Society of London 1903, 2 (1): 69-70 - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • D'angiolella, Annelise B.; Tony Gamble, Teresa C. S. Avila-Pires, Guarino R. Colli, Brice P. Noonan, and Laurie J. Vitt 2011. Anolis chrysolepis Duméril and Bibron, 1837 (Squamata: Iguanidae), Revisited: Molecular Phylogeny and Taxonomy of the Anolis chrysolepis Species Group. Bull. Mus. Comp. Zool. Harvard 160 (2): 35-63. - get paper here
  • Dirksen, L. & De la Riva, I. 1999. The lizards and amphisbaenians of Bolivia (Reptilia, Squamata): checklist, localities, and bibliography. Graellsia 55: 199-215
  • Griffin, L. E. 1917. A list of the South American lizards of the Carnegie Museum, with descriptions of four new species. Annals of the Carnegie Museum 11: 304-320. - get paper here
  • Guarnizo, Carlos E.; Fernanda P. Werneck, Lilian G. Giugliano, Marcella G. Santos, Jéssica Fenker, Lucas Sousa, Annelise B. D’Angiolella, Adriana R. dos Santos, Christine Strüssmann, Miguel T. Rodrigues, Tainá F. Dorado-Rodrigues, Tony Gamble, Guarin 2015. Cryptic lineages and diversification of an endemic anole lizard (Squamata, Dactyloidae) of the Cerrado hotspot. Molecular Phylogenetics and Evolution 94: 279-289, doi:10.1016/j.ympev.2015.09.005 - get paper here
  • Langstroth, Robert P. 2006. NOTAS SOBRE ANOLIS MERIDIONALIS BOETTGER, 1885 (SQUAMATA: IGUANIA: POLYCHROTIDAE) EN BOLIVIA Y COMENTARIOS SOBRE ANOLIS STEINBACHI. Kempffiana 2 (1): 154-172 - get paper here
  • Motte, Martha; Pier Cacciali 2009. Descripción de un neotipo para Anolis meridionalis Boettger, 1885 (Sauria: Polychrotidae). Cuadernos de Herpetología 23 (1):19-24 - get paper here
  • Nicholson, K.E. 2002. Phylogenetic analysis and a test of the current infrageneric classification of Norops (beta Anolis). Herpetological Monographs 16: 93-120 - get paper here
  • NICHOLSON, KIRSTEN E.; BRIAN I. CROTHER, CRAIG GUYER & JAY M. SAVAGE 2012. It is time for a new classification of anoles (Squamata: Dactyloidae). Zootaxa 3477: 1–108
  • Peters, James A. & Donoso-Barros, Roberto 1970. Catalogue of the Neotropical Squamata: Part II. Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 293 pp. - get paper here
  • Poe, S. 2004. Phylogeny of anoles. Herpetological Monographs 18: 37-89 - get paper here
  • Poe, S. 2013. 1986 Redux: New genera of anoles (Squamata: Dactyloidae) are unwarranted. Zootaxa 3626 (2): 295–299 - get paper here
  • RIBEIRO-JÚNIOR, MARCO A. 2015. Catalogue of distribution of lizards (Reptilia: Squamata) from the Brazilian Amazonia. I. Dactyloidae, Hoplocercidae, Iguanidae, Leiosauridae, Polychrotidae, Tropiduridae. Zootaxa 3983 (1): 001–110
  • Vanzolini,P.E. & Williams,E.E. 1970. South American anoles: the geographic differentiation and evolution of the Anolis chrysolepis species group (Sauria, Iguanidae). Arq. Zool. (São Paulo) 19: 1-298 - get paper here
 
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