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Atheris ceratophora WERNER, 1896

IUCN Red List - Atheris ceratophora - Vulnerable, VU
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Higher TaxaViperidae, Viperinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesG: Usambara-Buschviper, Gehörnte Buschviper 
SynonymAtheris ceratophora WERNER 1896: 194
Atheris ceratophorus — RASMUSSEN 1981: 181
Atheris ceratophorus — BROADLEY & HOWELL 1991: 23
Atheris ceratophorus — WELCH 1994: 19
Atheris ceratophora — BROADLEY 1998: 123
Atheris ceratophora — JOGER & COURAGE 1999
Atheris ceratophora — MCDIARMID, CAMPBELL & TOURÉ 1999: 354
Atheris ceratophorus — SPAWLS et al. 2001
Atheris ceratophora — WALLACH et al. 2014: 61
Atheris ceratophora — SPAWLS et al. 2018: 586 
DistributionTanzania (Usambara, Uluguru, Uzungwe Mountains), elevation 1400-3000 m.

Type locality: Usambara Mountains, Tanzania.  
Reproductionovovivparous 
TypesHolotype: BMNH 1946.1.18.23 
Diagnosis 
CommentVenomous!

Habitat: trees (arboreal)

Distribution: Has been erroneously listed for Benin but does not occur there (Hughes 2013). 
Etymologynamed after Greek “kéras” (horn) and “phéro” = bearing. 
References
  • Akeret, B. 2004. Grundlagen der Giftschlangenhaltung. Draco 5 (17): 37-53 - get paper here
  • Böhme,W. 1987. Seltene Reptilien aus dem Uzungwe-Gebirge, Süd-Tansania. Herpetofauna 9 (48): 27-34 - get paper here
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1998. A review of the genus Atheris Cope (Serpentes: Viperidae), with the description of a new species from Uganda. The Herpetological Journal 8: 117-135. - get paper here
  • Cimatti, E. 2006. Reptile Parks in Tanzania. Reptilia (GB) (48): 52-59 - get paper here
  • Dobiey, M. & Vogel, G. 2007. Venomous Snakes of Africa - Giftschlangen Afrikas. Edition Chimaira, Terralog 15, 150 pp. - get paper here
  • Emmrich D 1997. Further remarks on the rare Usambara Forest viper, Atheris ceratophora, WERNER 1895. Mitt. Zool. Mus. Berlin 73: 131-151 - get paper here
  • Ernst, R. & Rödel, M. O. 2002. A new Atheris species (Serpentes: Viperidae), from Taï National Park, Ivory Coast. The Herpetological Journal 12: 55-61 - get paper here
  • Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248
  • Hirschmann, W. 2008. Mehrjährige Haltung und erfolgreiche Vermehrung der Gehörnten Buschviper, Atheris ceratophora WERNER, 1896. Sauria 30 (3): 5-12 - get paper here
  • Jachan, G. 2010. Pflege und Vermehrung der Usambara-Buschviper Atheris ceratophora. Reptilia (Münster) 15 (83): 58-68 - get paper here
  • Joger,U. & Courage,K. 1999. Are palearctic ‘rattlesnakes’ (Echis and Cerastes) monophyletic? Kaupia (Darmstadt) (8): 65-81
  • Love, B. 2017. Action zeigen - ohne Fotos. Reptilia (Münster) 22 (128): 60-61 - get paper here
  • Loveridge,A. 1933. Reports on the scientific Results of an Expedition to the Southwestem Highlands of Tanganyika Territory. VII. Herpetology. Bull. Mus. Comp. Zool. Harvard 74: 197-415 - get paper here
  • Lyakurwa, J.V., Howell, K.M., Munishi, L.K., Treydte, A.C. 2019. Uzungwa Scarp Nature Forest Reserve; a unique hotspot for reptiles in Tanzania. Acta Herpetologica 14 (1): 3-14 - get paper here
  • Mallow, D. Ludwig, D. & Nilson, G. 2003. True Vipers: Natural History and Toxinology of Old World Vipers. Krieger, Malabar, Florida, 410 pp. [review in HR 35: 200, Reptilia 35: 74]
  • McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. [type catalogue] Herpetologists’ League, 511 pp.
  • Menegon, M.; Loader, S.P.; Marsden, S.J.; Branch, W.R.; Davenport, T.R.B. & Ursenbacher, S. 2014. The genus Atheris (Serpentes: Viperidae) in East Africa: Phylogeny and the role of rifting and climate in shaping the current pattern of species diversity. Molecular Phylogenetics and Evolution 79: 12-22 - get paper here
  • Menegon, Michele; Nike Doggart, Nisha Owen 2008. The Nguru mountains of Tanzania, an outstanding hotspot of herpetofaunal diversity. Acta Herpetologica 3 (2): 107-127
  • MENEGON, MICHELE; TIM R.B. DAVENPORT & KIM M. HOWELL 2011. Description of a new and critically endangered species of Atheris (Serpentes: Viperidae) from the Southern Highlands of Tanzania, with an overview of the country’s tree viper fauna. Zootaxa 3120: 43–54 - get paper here
  • O’Shea, M. 2018. The Book of Snakes. Ivy Press / Quarto Publishing, London, - get paper here
  • Phelps, T. 2010. Old World Vipers. Edition Chimaira, Frankfurt, 558 pp. [critical review in Sauria 33 (3): 19 and HR 43: 503]
  • Rasmussen, J.B. & K.M. HOWELL 1982. The current status of the rare Usambara mountain forest-viper, Atheris ceratophorus Werner, 1895, including a probable new record of Atheris nitschei rungweensis Bogert, 1940, and a discussion of its validity (Reptilia, Serpentes, Viperidae). Amphibia-Reptilia 3: 269-277 - get paper here
  • Rasmussen,J.B. 1981. The snakes from the rainforest of the Usambara Mountains, Tanzania: a checklist and key. Salamandra 17 (3-4): 173-188 - get paper here
  • Rovero, F., Menegon, M., Fjeldså, J., Collett, L., Doggart, N., Leonard, C., Norton, G., Owen, N., Perkin, A., Spitale, D., Ahrends, A., Burgess, N. D. 2014. Targeted vertebrate surveys enhance the faunal importance and improve explanatory models within the Eastern Arc Mountains of Kenya and Tanzania. Diversity and Distributions. doi: 10.1111/ddi.12246 - get paper here
  • Spawls, S., Branch, B. 1995. The Dangerous Snakes of Africa. Blandford, London, 192 pp.
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Spawls, Steve; Kim Howell, Harald Hinkel, Michele Menegon 2018. Field Guide to East African Reptiles. Bloomsbury, 624 pp. - get paper here
  • Spranger, Tade Matthias 2012. Haltung und Handel „exotischer” Tiere als Gegenstand des Polizei- und Ordnungsrechts. Draco 13 (51): 44-53 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Werner, F. 1895. Über einige Reptilien aus Usambara (Deutsch-Ostafrika). Verhandlungen der Kaiserlich-Königlichen Zoologisch-Botanischen Gesellschaft in Wien, 45: 190—194 - get paper here
  • Werning, H. & Wolf, C. 2007. Gebirge und Hochland - Reptilien und Amphibien in einem extremen Lebensraum. Draco 7 (27): 4-27 - get paper here
 
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