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Atractus caxiuana DA COSTA PRUDENTE & SANTOS-COSTA, 2006

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common Names 
SynonymAtractus caxiuana DA COSTA PRUDENTE & SANTOS-COSTA 2006
Atractus caxiuana — WALLACH et al. 2014: 70 
DistributionS Brazil (Pará) through Colombia (Vaupés)

Type locality: “Estação Científica Ferreira Penna”, Eastern Amazonia, Pará State, Brazil Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesHolotype: MPEG 19657 
CommentDiagnosis: 17-17-17 smooth dorsal scales without pits; 162 ventrals and 30 pairs of subcaudals in the single measurable male and 171 ventrals and 24 pairs of subcaudals in the single avaliable female; the three available specimens (two males and one female) has frontal wider than long; nasal contacting loreal, internasals, rostral, and the two first pairs of supralabials; loreal contacting internasal, and second and third supralabials; preocular absent; two postoculars; two light spots on the back of the head; wide-black median ventral band; and subcaudals totally black.

Diagnosis. Atractus caxiuana is distinguished from all other congeners by the following combination of characters: loreal contacting the internasal (but see below); 17/17/17 dorsal scale rows with paired apical pits in both sexes and supra-anal tubercles present only in males; one or two postoculars; loreal moderately long; temporals 1+2; seven supralabials, third and fourth contacting orbit; seven infralabials, first three contacting chinshields; five or six maxillary teeth; four gular scale rows; four preventrals; 171–184 ventrals in females, 160– 162 in males; 18–24 subcaudals in females, 25–30 in males; dorsum mostly dark brown, except for dispersed beige blotches on snout, a creamy incomplete collar covering the parietal, occipital and temporal regions of the head, and two pale brown longitudinal stripes covering the centre of first two dorsal scale rows of the body; belly creamy with lateral portion of the ventral scales dark brown, forming paraventral stripes; small body size, females reaching 265 mm SVL, males 261 mm SVL; small tail length in females (9.0–9.2% SVL), moderately long in males (9.2– 13.2% SVL); hemipenis slightly bilobed, non-capitate and non-calyculate [Passos et al. 2013: 40].

Habitat: rainforest 
References
  • Passos, P., R. Fernandes and Borges-Nojosa, D.M. 2007. A New Species of Atractus (Serpentes: Dipsadinae) from a Relictual Forest in Northeastern Brazil. Copeia 2007 (4): 788–797 - get paper here
  • PASSOS, PAULO; LUCIANA O. RAMOS, PEDRO H. PINNA, ANA L.C. PRUDENTE 2013. Morphological variation and affinities of the poorly known snake Atractus caxiuana (Serpentes: Dipsadidae). Zootaxa 3745 (1): 035–048 - get paper here
  • Prudente, A. L. da & Santos-Costa, M. C. dos 2006. A new species of Atractus Wagler, 1828 (Serpentes: Colubridae) from Eastern Amazonia, Brazil. Zootaxa 1285: 21–29 - get paper here
  • Prudente, Ana L.C and Paulo Passos 2008. New Species of Atractus Wagler, 1828 (Serpentes: Dipsadinae) from Guyana Plateau in Northern Brazil. Journal of Herpetology 42 (4): 723 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
 
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