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Atractus pyroni ARTEAGA, MEBERT, VALENCIA, CISNEROS-HEREDIA, PEÑAFIEL, REYES-PUIG, VIEIRA-FERNANDES & GUAYASAMIN, 2017

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesE: Pyron’s Groundsnake
S: Tierrera de Pyron 
SynonymAtractus pyroni ARTEAGA, MEBERT, VALENCIA, CISNEROS-HEREDIA, PEÑAFIEL, REYES-PUIG, VIEIRA-FERNANDES & GUAYASAMIN 2017 
DistributionEcuador

Type locality: between Balzapamba and Bilován, province of Bolívar, Ecuador (S1.83601, W79.13322; 2026 m elevation).  
Reproductionoviparous 
TypesHolotype. MZUTI 5107 (Fig. 7), adult male collected by José L. Vieira-Fernandes and Carlos Durán on May 23, 2016. 
CommentDiagnosis. Atractus pyroni is placed in the genus Atractus as diagnosed by Savage (1960), based on phylogenetic (Fig. 1) and morphological (Table 3) evidence. It is included in the A. roulei group due to its 15/15/15 dorsal scale rows and its phylogenetic position (Fig. 1). The species is diagnosed based on the following combination of characters: (1) 15/15/15 smooth dorsals; (2) one postocular; (3) loreal long; (4) temporals 1+2; (5) six supralabials, third and fourth contacting orbit; (6) five infralabials, first four contacting chinshields (7) eight maxillary teeth; (8) 3 gular scale rows; (9) 2 preventrals; (10) 143 ventrals; (11) 16 subcaudals; (12) dorsal ground color dark brown with a series of light golden brown paravertebral scales running along the entire dorsum (Fig. 7); (13) venter dark brown with scattered scales of a lighter color; (14) 443 mm SVL; (15) 34 mm TL.

Comparisons. Atractus pyroni is compared to members of the A. roulei species group: A. carrioni, A. lehmanni, and A. roulei (Fig. 2). From A. carrioni, it differs by having a loreal. From A. lehmanni and A. roulei, it differs in size and color pattern. Atractus pyroni is 443 mm in SVL; whereas A. lehmanni is 262–321 in SVL, and A. roulei is 230–396. Both A. lehmanni and A. roulei have uniform dorsal ground color, whereas A. pyroni has a distinct dorsal bicolored pattern (Fig. 7). Finally, in life, A. pyroni is darker than the remaining members of the A. roulei species group and has a ventral pattern that, instead of having fine speckles, has conspicuous scattered blotches of a contrasting color.

Conservation status: Data Deficient 
EtymologyNamed after R. Alexander Pyron, one of the most prolific contemporary herpetologists, in recognition of his invaluable contribution to systematics and evolution of the world’s reptiles. 
References
  • Arteaga A, Mebert K, Valencia JH, Cisneros-Heredia DF, Peñafiel N, Reyes-Puig C, Vieira-Fernandes JL, Guayasamin JM 2017. Molecular phylogeny of Atractus (Serpentes, Dipsadidae), with emphasis on Ecuadorian species and the description of three new taxa. ZooKeys 661: 91-123. - get paper here
 
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