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Atractus reticulatus (BOULENGER, 1885)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
SubspeciesAtractus reticulatus scrocchii ALVAREZ, REY & CEI 1992
Atractus reticulatus reticulatus (BOULENGER 1885) 
Common NamesReticulate Ground Snake 
SynonymGeophis reticulatus BOULENGER 1885: 87
Geophis reticulatus — BOULENGER 1886: 430
Atractus reticulatus — BOULENGER 1894: 311
Atractus reticulatus — PETERS et al. 1970: 33
Atractus reticulatus scrocchii ALVAREZ et al. 1992
Atractus reticulatus — PASSOS et al. 2005
Atractus reticulatus — WALLACH et al. 2014: 79 
DistributionParaguay, S Brazil (Rio Grande do Sul, Paraná, São Paulo), Argentina (Corrientes, Misiones ?), Uruguay [HR 35: 189]
Colombia [Castro,F. (pers. comm.)]

Type locality: São Lourenço, Estado do Rio Grande do Sul, Brazil.  
TypesHolotype: BMNH 1946.1.2.7
Holotype: UNNE-CHC 00256, collected by B.B. Alvarez and L. Rey 28 Dec 1991. [scrocchii] 
DiagnosisDiagnosis: Atractus reticulatus is distinguished from all congeners by the following combination of characters: (1) 15/15/15 smooth dorsal scale rows; (2) two postoculars; (3) loreal moderate; (4) temporals 1+2; (5) seven supralabials, third and fourth contacting orbit; (6) six or seven infralabials, first four contacting chinshields; (7) six to eight maxillary teeth; (8) usually three gular scale rows; (9) usually three preventrals; (10) 149–165 ventrals in females, 130–151 in males; (11) 19–30 subcaudals in females, 24–34 in males; (12) dorsum ground colour dark grayish brown or black, reticulating with pale colour; (13) venter immaculate creamish white; (14) moderate body size, females reaching 390 mm SVL, males 279 mm SVL; (15) tail short to moderate in females (7.8–13.5% SVL), short to long (7.9–17.1% SVL) in males; (16) moderately bilobed, semicapitate, and semicalyculate hemipenis [from Passos et al. 2010]. 
  • Alvarez B B. Rey L. Cei J M. 1992. A new subspecies of the Reticulatus group, genus Atractus, from southeastern South America (Serpentes, Colubridae). MUSEO REGIONALE DI SCIENZE NATURALI BOLLETTINO (TURIN). 10 (2): 249-256.
  • Bérnils, R.S.; Batista, M.A. & Bertelli, P.W. 2001. Cobras e lagartos do Vale: levantamento das espécies de Squamata (Reptilia, Lepidosauria) da Bacia do Rio Itajai, Santa Catarina, Brasil. Rev. Est. Ambientais (Blumenau) 3 (1): 69-79
  • Boulenger, G. A. 1885. Second list of reptiles and batrachians from the province Rio Grande do Sul, sent to the Natural History Museum by Dr. H. van Ihering. Ann. Mag. nat. Hist. (5) 16: 85-88 - get paper here
  • Boulenger, G.A. 1886. A synopsis of the reptiles and batrachians of the province Rio Grande do Sul, Brazil. Ann. Mag. nat. Hist. (5) 18: 423-445 - get paper here
  • Boulenger, George A. 1894. Catalogue of the snakes in the British Museum (Natural History). Volume II., Containing the Conclusion of the Colubridæ Aglyphæ. British Mus. (Nat. Hist.), London, xi, 382 pp. - get paper here
  • Cacciali, P.; Villalba, R. & Yanosky, A.A. 2007. A NEW SPECIES OF ATRACTUS (SERPENTES:COLUBRIDAE: DIPSADINAE) FROM THE ATLANTIC FOREST OF ALTO PARANÁ, PARAGUAY. South American J. Herp. 2 (2): 83-88 - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • Carreira, S.; Brazeiro, A.; Camargo, A.; da Rosa, I.; Canavero, A. & Maneyro, M. 2012. DIVERSITY OF REPTILES OF URUGUAY: KNOWLEDGE AND INFORMATION GAPS. Bol. Soc. Zool. Uruguay (2a época) 21 (1-2): 9-29 - get paper here
  • Carreira, Santiago; Ignacio Lombardo & Federico Achaval-Coppes. 2012. NUEVOS REGISTROS DE OFIDIOS POCO CONOCIDOS EN URUGUAY (REPTILIA: SQUAMATA: SERPENTES). Bol. Soc. Zool. Uruguay (2a época) 21 (1-2): 70-74 - get paper here
  • Lucchesi Balestrin, R. & Di-Bernardo, M. 2005. Reproductive biology of Atractus reticulatus (BOULENGER 1885) (Serpentes, Colubridae) in Southern Brazil. The Herpetological Journal 15: 195-199
  • Outeiral, Arlete B.; Rafael L. Balestrin, Lize H. Cappellari, Thales de Lema, Vanda Lúcia Ferreira 2018. Snake assemblage from Serra do Sudeste, Pampa Biome in Southern Brazil Herpetology Notes 11: 733-745 - get paper here
  • Passos, P., R. Fernandes and Borges-Nojosa, D.M. 2007. A New Species of Atractus (Serpentes: Dipsadinae) from a Relictual Forest in Northeastern Brazil. Copeia 2007 (4): 788–797 - get paper here
  • Passos, P., R. Fernandes and N. Zanella 2005. A new species of Atractus (Serpentes: Colubridae) from Southern Brazil. Herpetologica 61 (2): 209-218 - get paper here
  • Passos, P.; Fernandes, R.; Bernils, R.S. & Moura-Leite, J.C. de 2010. Taxonomic revision of the Brazilian Atlantic Forest Atractus (Reptilia: Serpentes: Dipsadidae). Zootaxa 2364: 1–63 - get paper here
  • Passos, Paulo; Altagratia Chiesse, Omar Torres-Carvajal, and Jay M. Savage 2009. TESTING SPECIES BOUNDARIES WITHIN THE ATRACTUS OCCIPITOALBUS COMPLEX (SERPENTES: DIPSADIDAE). Herpetologica 65 (4): 384-403 - get paper here
  • Pérez-Santos,C. & Moreno, A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Peters, James A.; Donoso-Barros, Roberto & Orejas-Miranda, Braulio 1970. Catalogue of the Neotropical Squamata: Part I Snakes. Part II Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 347 pp. - get paper here
  • Pizzatto, Lígia; Maurício Cantor, Juliana Lima de Oliveira, Otavio A.V. Marques, Vinicius Capovilla, and Marcio Martins 2008. Reproductive Ecology of Dipsadine Snakes, With Emphasis on South American Species. Herpetologica 64 (2): 168-179 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zaracho, Víctor Hugo; Ingaramo, María del Rosario; Semhan, Romina Valeria; Etchepare, Eduardo; Acosta, José Luis; Falcione, Ana Camila; Álvarez, Blanca 2014. Herpetofauna de la Reserva Natural Provincial Isla Apipé Grande (Corrientes, Argentina) Cuad. herpetol. 28 (2): 153-160 - get paper here
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