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Atractus werneri PERACCA, 1914

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesWerner's Ground Snake 
SynonymAtractus werneri PERACCA 1914: 102
Atractus werneri — DUNN 1944: 21
Atractus werneri — PASSOS & LYNCH 2010
Atractus werneri — WALLACH et al. 2014: 84 
DistributionColombia (Cundinamarca: Western slopes of Eastern Cordillera), elevation 1200-1800 m

Type locality: Viotá, Dep. de Cundinamarca, Colombia, 1830 m elevation, (04° 26’ N, 74° 31’ W).  
Reproductionoviparous 
TypesHolotype: MNHN 91.506 
DiagnosisDiagnosis.—Atractus werneri is distinguished from all congeners by having: (1) 17/ 17/17 smooth dorsal scale rows; (2) two postoculars; (3) loreal long; (4) temporals 1 + 2; (5) generally seven supralabials, third and fourth contacting orbit; (6) generally seven infralabials, first three contacting chinshields; (7) 6–9 maxillary teeth; (8) three or four gular scale rows; (9) three or four preventrals; (10) 158–174 ventrals in females, 148–160 in males; (11) 21–36 subcaudals in females, 27– 37 in males; (12) dorsum uniformly brown or having conspicuous vertebral and dorsolateral lines; (13) venter predominantly creamish white, occasionally having diffuse and irregu- lar dark brown blotches; (14) moderate body size, females reaching 346 mm SVL, males 330 mm SVL; (15) small to moderate tail length in females (8.7–15.2% SVL), moderate to long in males (13.2–17.1% SVL); (16) hemipenis slightly bilobed, semicapitate, and semicalyculate. 
CommentThe date of publication of Peracca’s paper is usually given as 1914. However, it has been shown that his paper was actually published at least in early 1913 but most likely in 1912.

Similar species: A. obtusirostris and A. melanogaster. 
EtymologyNamed after Franz Werner (1867-1939), Austrian zoologist. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Dunn, Emmett R. 1944. Herpetology of the Bogota area. Revista de la Academia Colombiana de Ciencias Exacta, Fisicas y Naturales 6: 68-81
  • Passos, Paulo and John D. Lynch 2010. Revision of Atractus (Serpentes: Dipsadidae) from Middle and Upper Magdalena Drainage of Colombia. Herpetological Monographs 24 (1): 149-173 - get paper here
  • Peracca, M.G. 1914. Reptiles et Batraciens de Colombie. Pp. 96-111 In: Fuhrmann, O. and E. Mayor (eds.), Voyage d'exploration scientifique en Colombie. Memoires de la Societe neuchateloise des Sciences naturelles, 5
  • Pérez-Santos,C. & Moreno, A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
 
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