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Bachia marcelae DONOSO-BARROS & GARRIDO, 1964

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Higher TaxaGymnophthalmidae (Cercosaurinae, Bachiinae), Sauria, Gymnophthalmoidea, Squamata (lizards)
Subspecies 
Common NamesE: Marcela’s Bachia, Marcela’s Worm lizard 
SynonymBachia marcelae DONOSO-BARROS & GARRIDO 1964: 3
Bachia lineata marcelae — DONOSO-BARROS 1968: 118
Bachia heteropa marcelae – DIXON 1973: 33
Bachia heteropa marcelae — JOHN et al. 2012
Bachia marcelae — MURPHY et al. 2019 
DistributionVenezuela

Type locality: “Bosque tropical La Luz, Edo. Barinas, Venezuela.”, given as La Luz, Edo. Barinas, Venezuela (8° 23′ N, 69° 49′ W) by Murphy et al. 2019.  
Reproductionoviparous 
TypesHolotype: MZUC 8578 (Concepción, Chile), male, fide Murphy et al. 2019, was: DB 6312061, i.e. private collection of Donoso-Barros, collected on 12 December 1963 by R. Donoso-Barros and A. Garrido. Paratype: female, probably also in MZUC. 
DiagnosisDiagnosis A Bachia with (1) no prefrontal scales; (2) two supraoculars; (3) the frontonasal is hexagonal; (4) four digits are present on the manus and two on the pes (5) a band of imbricate hexagonal scales on the dorsum; (6) scales around mid-body 25; (7) 39–42 transverse rows of dorsals; (8) the fifth upper labial contacts primary temporal; (9) the gulars are in five rows; (10) the second pair of chin shield are in medial contact; (11) a sub-rectangular interparietal present (from Murphy et al. 2019). 
CommentSynonymy partly that of Dixon (1973) who synonymized B. marcelae with B. heteropa. However, more recent authors recognized marcelae as valid. 
References
  • Bauer, A.M. and R. Günther 1994. An annotated type catalogue of the teiid and microteiid lizards in the Zoological Museum, Berlin (Reptilia: Squamata: Teiidae and Gymnophthalmidae). Mitt. Zool. Mus. Berlin 70:267-280 - get paper here
  • Bentz, Ehren J.; Mel José Rivera Rodríguez, Rebecca R. John, Robert W. Henderson, and Robert Powell 2011. Population Densities, Activity, Microhabitats, and Thermal Biology of a Unique Crevice- and Litter-dwelling Assemblage of Reptiles on Union Island, St. Vincent and the Grenadines. Herp. Cons. Biol. 6 (1): 40-50 - get paper here
  • Boos, H.E.A. 1984. The terrestrial reptiles of Monos island. Living World - get paper here
  • Daan, S. & Hillenius,D. 1966. Catalogue of the type specimens of amphibians and reptiles in the Zoological Museum, Amsterdam. Beaufortia 13: 117-144
  • Daudin, Jacques; de Silva, Mark. 2007. An annotated checklist of the amphibians and terrestrial reptiles of the Grenadines with notes on their local natural history and conservation. Applied Herpetology 4 (2):163-175 - get paper here
  • Disi, Ahmad; Zuhair S. Amr, Nashat Hamidan 2014. Diversity, Threats, and Conservation of the Terrestrial and Freshwater Herpetofauna of Jordan. Russ. J. Herpetol. 21 (3): 221-233 - get paper here
  • Dixon, J.R. 1973. A systematic review of the teiid lizards, genus Bachia, with remarks on Heterodactylus and Anotosaura. Misc. Publ. Univ. Kans. Mus. Nat. Hist. 57: 1-47 - get paper here
  • Donoso-Barros, R. 1968. The lizards of Venezuela (checklist and key). Carib. J. Sci. 8 (3-4): 105-122
  • Donoso-Barros, R. and A. Garrido R. 1964. Nuevo Teiidae de Venezuela, Bachia marcelae, nov. sp. Publ. Ocas. Mus. Cienc. Nat., Caracas (Zool.) 8: 1-7
  • Esqueda, Luis Felipe, Enrique La Marca, Marco Natera and Pietro Battiston 2001. Noteworthy reptilian state records and a lizard species new to the herpetofauna of Venezuela. Herpetological Review 32 (3):198-200 - get paper here
  • Gorzula, Stefan & Senaris, J. Celsa 1999. In: Contribution to the herpetofauna of the Venezuelan Guayana. I: a data base. Scientia Guaianae, Caracas, No. 8 [1998], 269+ pp.; ISBN 980-6020-48-0
  • John, Rebecca R., Ehren J. Bentz, Mel Jose Rivera Rodriguez, Aaron M. Bauer and Robert Powell. 2012. Bachia heteropa. Catalogue of American Amphibians and Reptiles (894): 1-9 - get paper here
  • Kramer, Eugen 1979. Typenkatalog der Echsen im Naturhistorischen Museum Basel (BM), Stand 1978. [type catalogue] Revue Suisse de Zoologie 86 (1): 159-166 - get paper here
  • Markezich, A.L. 2002. New distribution records of reptiles from Western Venezuela. Herpetological Review 33 (1): 69-74 - get paper here
  • Mijares-Urrutia, A. & Arends R., Alexis 1999. Additional New Regional and Local Records of Amphibians and Reptiles from the State of Falcón, Venezuela. Herpetological Review 30 (2): 115 - get paper here
  • Murphy, John C.; Daniele Salvi, Joana L. Santos, Alvin L. Braswell, Stevland P. Charles, Amaél Borzée & Michael J. Jowers 2019. The reduced limbed lizards of the genus Bachia (Reptilia, Squamata, Gymnophthalmidae); biogeography, cryptic diversity, and morphological convergence in the eastern Caribbean. Organisms Diversity & Evolution - get paper here
  • Powell, Robert and Robert W. Henderson 2017. Notes on the diets of some amphibians and reptiles of the St. Vincent and Grenada Banks (Lesser Antilles). IRCF Reptiles & Amphibians 24 (2): 135–138 - get paper here
  • Ribeiro-Júnior, Marco A.; Marcélia B. da Silva, and Jucivaldo D. Lima 2016. A New Species of Bachia Gray 1845 (Squamata: Gymnophthalmidae) from the Eastern Guiana Shield. Herpetologica 72 (2): 148-156. - get paper here
  • Rivas Fuenmayor, G.; Ugueto, G.; Rivero, R. & Miralles, A. 2005. The Herpetofauna of Isla de Margarita, Venezuela: New Records and Comments. Carib. J. Sci. 41 (2): 346-351
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Schwartz, A. & Henderson, R.W. 1991. Amphibians and Reptiles of the West Indies. University of Florida Press, Gainesville, 720 pp.
  • Señaris, J. Celsa; María Matilde Aristeguieta Padrón, Haidy Rojas Gil y Fernando J. M. Rojas-Runjaic 2018. Guía ilustrada de los anfibios y reptiles del valle de Caracas, Venezuela. Ediciones IVIC, Instituto Venezolano de Investigaciones Científicas (IVIC). Caracas, Venezuela. 348 pp.
  • Vanzolini,P.E. 1961. Bachia: Especies Brasileiras e conceito generico (Sauria: Teiidae). Papéis Avulsos Zoologia (São Paulo) 14: 193-209 - get paper here
 
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