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Barisia imbricata (WIEGMANN, 1828)

IUCN Red List - Barisia imbricata - Least Concern, LC
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Higher TaxaAnguidae (Gerrhonotinae), Diploglossa, Anguimorpha, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Imbricate Alligator Lizard, Transvolcanic Alligator Lizard
S: Escorpión de Transvolcánico 
SynonymGerrhonotus imbricatus WIEGMANN 1828: 379
Gerrhonotus imbricatus — DUMÉRIL & BIBRON 1839: 407
Barissia imbricata — GRAY 1845: 55
Barissia imbricata — COPE 1878: 97
Gerrhonotus (Barissia) imbricatus — BOCOURT 1878: 363
Gerrhonotus imbricatus — BOULENGER 1885: 272
Barisia imbricata imbricata — TIHEN 1949: 240
Gerrhonotus lichenigerus WAGLER 1833 (1830 fide GOOD et al. 1993)
Gerrhonotus adspersus WIEGMANN 1834
Gerrhonotus Lichenigerus — DUMÉRIL & BIBRON 1839: 408
Gerrhonotus olivaceus BAIRD 1859
Gerrhonotus olivaceus — O’SHAUGHNESSY 1873: 48
Barisia imbricata imbricata — SMITH & TAYLOR 1950: 201
Gerrhonotus imbricatus — WERMUTH 1969: 18
Barisia imbricata — LINER 1994
Barisia imbricata — ZALDIVAR-RIVERON et al. 2005
Barisia imbricata — CANSECO-MÁRQUEZ & GUTIÉRREZ-MAYÉN 2010 
DistributionMexico (Jalisco, Michoacán, Morelos, Puebla, Oaxaca, Veracruz, Hidalgo, Guanajuato, Distrito Federal, Chihuahua [HR 31: 112], Zacatecas, Aguascalientes, Guerrero, Nayarit).

Type locality: Mexico; restricted to Distrito Federal by SMITH & TAYLOR 1950. Elevation 9,800-13,000 feet.  
Reproductionovovivparous 
TypesHolotype: ZMB 1156 (imbricata)
Holotype: ZSM 582/0 [lichenigerus]
Holotype: UMMZ 112704 [jonesi]
Syntype: USNM 3096 [Gerrhonotus olivaceus] 
DiagnosisDiagnosis (genus): Good 1988: 78 
CommentSubspecies: subspecies of B. imbricata have been elevated to full species.

Map: GUILLETTE & SMITH 1982: 24

Distribution: Not listed for San Luis Potosí by Lemos-Espinal & Dixon 2013. Not in Nuevo Leon fide LEMOS-ESPINAL et al. 2016. Not listed for San Luis Potosí by Lemos-Espinal et al. 2018 (checklist SLP). Not in Durango fide Lemos-Espinal (2018). Not in Sinaloa (Lemos-Espinal & Smith 2020). Reports from Quéretaro are now assigned to B. ciliaris, hence the species does not occur in Quéretaro fide Tepos-Ramírez et al. 2023.

Type species: Gerrhonotus imbricatus WIEGMANN 1828 is the type species of the genus Barisia GRAY 1838.

Definition (genus Barisia): Gerrhonotine lizards with the skull not widened or depressed ; frontal bone not in contact with the maxillae ; pterygoid teeth absent or vestigial; dorsal osteoderms with a well-defined, strongly thickened basal area; lateral fold moderately to well developed, with a moderate or large granular area ; sides of the neck finely granular; postero-ventral surfaces of the forearms and shanks granular; minimum number of scales in a single row across the nape six to ten; anterior internasals present; postrostral present or absent; subocular and postocular series well differentiated from each other; the suboculars extend to the lowest primary temporal; twelve to fourteen ventral scale rows (fide TIHEN 1949). 
EtymologyThe name imbricata is derived from the Latin imbricatus (overlapping, like roofing tiles and shingles) and refers to the overlapping dorsal scales.

The generic name was derived from the Greek word barys, meaning "heavy," apparently in reference to the robust body of B. imbricata (Lemos-Espinal & Dixon 2013). 
References
  • ARZATE-GARAY, LAURA S.; JOSELE R. FLORES-SANTIN, G. PAMELA HERNANDEZ-LOPEZ & CHARLES M. WATSON. 2022. BARISIA IMBRICATA (Transvolcanic Alligator Lizard). ARBOREAL HABITAT USE. Herpetological Review 53 (2): 320.
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  • Bocourt, M. E. 1873. In: A. Duméril, M. F. Bocourt, and F. Mocquard, (1870-1909), Etudes sur les reptiles, p. i-xiv; In Recherches Zoologiques pour servir a l'Histoire de Ia Faune de l'Amérique Centrale et du Mexique. Mission Scientifique au Mexique et dans l'Amérique Ce Imprimerie Impériale, Paris, Livr. 2-15, pp. 33-860. - get paper here
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  • Good, David A.; Bauer, Aaron M.;Günther, Rainer 1993. An Annotated Type Catalogue of the Anguimorph Lizards (Squamata: Anguidae, Helodermatidae, Varanidae, Xenosauridae) in the Zoological Museum, Berlin. Mitt. Zool. Mus. Berl. 69 (1): 45-56 - get paper here
  • Goyenechea Mayer-Goyenechea, Irene & Martha Gual-Díaz 2014. Reptilies en el Bosques mesófilos de Montaña de México. in: Gual-Diaz & Rendón-Correa, Bosques mesófilos de Montaña de México, CONABIO, pp. 263-278 - get paper here
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  • Guillette, L.J., Jr. & G. Casas-Andreu 1987. The Reproductive Biology of the High Elevation Mexican Lizard Barisia imbricata Herpetologica 43 (1): 29-38. - get paper here
  • Illescas-Aparicio, M., Clark-Tapia, R., González-Hernández, A., Vásquez-Díaz, Pedro R. & Aguirre-Hidalgo, V. 2016. Diversidad y riqueza herpetofaunística asociada al bosque de manejo forestal y áreas de cultivo, en Ixtlán de Juárez, Oaxaca. Acta Zoológica Mexicana (n.s.) 32(3): 359-369 - get paper here
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