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Bothriechis khwargi ARTEAGA, PYRON, BATISTA, VIEIRA, MENESES-PELAYO, SMITH, BARRIO-AMORÓS, KOCH, AGNE, VALENCIA, BUSTAMANTE & HARRIS, 2024

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Higher TaxaViperidae, Crotalinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Khwarg’s Eyelash-Pitviper.
G: Khwargs Lanzenotter
S: Víbora de pestañas de Khwarg 
SynonymBothriechis khwargi ARTEAGA, PYRON, BATISTA, VIEIRA, MENESES-PELAYO, SMITH, BARRIO-AMORÓS, KOCH, AGNE, VALENCIA, BUSTAMANTE, HARRIS 2024: 39 
DistributionColombia (Cordillera Oriental)

Type locality: Finca la Esperanza, vereda Sogamoso, municipio de Betulia, Santander department, Colombia (7.00600, -73.421608; 1195 m elevation.  
Reproduction 
TypesHolotype: UIS-R 4294 (Figs 17, 18b), adult female collected by Elson Meneses-Pelayo on November 15, 2017.
Paratypes: All labeled Bothriechis khwargi sp. nov. in Suppl. material 1. 
DiagnosisDiagnosis: Bothriechis khwargi sp. nov. is diagnosed based on the following combination of characters: (1) two triangular and moderately raised supraciliary scales; (2) anterior dorsal head scales keeled; (3) gular scales much smaller than chinshields; (4) 7–14 interoculolabials; (5) 3–4 canthals, some of which with raised triangular projections; (6) loreal in contact with preocular in ~1/3 of specimens; (7) yellow morph absent; (8) dorsal bands absent, faint, or restricted to top of dorsum in juveniles; (9) opposing kidney shaped dorsal marks absent; (10) black speckles on dorsal scales absent; (11) black speckling on ventral surfaces absent; (12) ventral surfaces entirely white in most individuals; (13) iris pale
green or straw yellow; (14) 21–23 dorsal scale rows at mid-body; (15) 144–153 ventrals in males, 145–154 in females; (16) maximum total length in males 219 mm, in females 610 mm. (Arteaga et al. 2024: 39) 
CommentVenomous!

Synonymy: Reyes-Velasco 2024 argued that B. torvus, B. khwargi, B. klebbai, B. rasikusumorum, B. rahimi, B. nitidus, B. torvus, and B. hussaini should be synonymized with B. schlegelii, given their genetic and morphological similarity. Reyes-Velasco 2024 thus argued that only B. nigroadspersus, B. schlegelii, and B. supraciliaris should be maintained. We agree that the former 5 species are not very much differentiated from B. schlegelii and should be considered preliminary species. We have not adopted Reyes-Velasco’s revision yet as there are reportedly other studies in the process of publication, hence the taxonomy of Bothriechis is in flux and more changes are likely to come soon.

Sympatry: B. klebbai 
References
  • Arteaga A, Pyron RA, Batista A, Vieira J, Meneses Pelayo E, Smith EN, Barrio Amorós CL, Koch C, Agne S, Valencia JH, Bustamante L, Harris KJ 2024. Systematic revision of the Eyelash Palm-Pitviper Bothriechis schlegelii (Serpentes, Viperidae), with the description of five new species and revalidation of three. Evolutionary Systematics 8(1): 15-64 - get paper here
  • Reyes-Velasco, J. 2024. A revision of recent taxonomic changes to the eyelash palm pitviper, Bothriechis schlegelii (Serpentes, Viperidae). Herpetozoa, 37: 305-318 - get paper here
 
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