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Bronchocela jubata DUMÉRIL & BIBRON, 1837

IUCN Red List - Bronchocela jubata - Least Concern, LC
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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Maned forest lizard 
SynonymBronchocela jubata DUMÉRIL & BIBRON 1837: 397
Bronchocele [sic] intermedia BERTHOLD 1842 (non Calotes intermedia PETERS & DORIA, 1878)
Calotes (Bronchocele) intermedius BERTHOLD 1842 (fide BÖHME 2012: 103
Calotes intermedius — BERTHOLD 1846: 12
Calotes jubatus — BOULENGER 1885: 318
Calotes jubatus — DE ROOIJ 1915: 123
Calotes jubatus — SMITH 1935: 185
Bronchocela jubata — MOODY 1980
Bronchocela petersidoriai BOHME & BlSCHOFF 1984 (nomen novum)
Bronchocela jubata — MANTHEY & SCHUSTER 1999: 28
Bronchocela jubata — PIANKA & VITT 2003: 151
Broncochela jubata — JANIAWATI et al. 2016 (in error) 
DistributionIndonesia (Singkep, Java, Bali, Celebes = Sulawesi, Karakelang, Salibabu; Nias Island, Singkap Island, Borneo: Kalimantan [DAS, pers. comm. to J.Hallermann),
Philippines (Mindanao: ZMB 16305),
India (questionable - see comment),
Cambodia [see comment]
SC Thailand

intermedia: Type locality: Sunda Islands restricted to Java.

Type locality: Pondichery (South India) and Java, restricted by Smith (1935 ) to Java. Lectotype locality: Java, Indonesia  
Reproductionoviparous 
TypesLectotype: MNHN-RA 2542, adult male, designated by Amarasinghe et al. 2022. Paralectotypes: MNHN-RA 2541 and (ii) MNHN-RA 2541A (both lost) and MNHN-RA 2543; Other specimens: RMNH, ZFMK, MZB, ZMH, ZMUC, TCWC, ZMB, UIMZ 
DiagnosisDiagnosis: A species of Bronchocela inhabiting the Greater Sunda Islands (Sumatra, Java, Borneo, and Bali), characterized as follows: morphologically most similar to its allopatric congener on Sumatra Island, B. hayeki (see Amarasinghe et al. 2022) and sympatric congener, B. cristatella, but differs by having a comparatively larger gular sac (smaller in B. hayeki and B. cristatella), ventral scales arranged in 10–12 non-enlarged rows (vs. 8–10 enlarged rows in B. hayeki and 10–14 slightly-enlarged rows in B. cristatella), lower number of mid body scale rows, 33–59 (vs. 64–75 in B. hayeki and 50–106 in B. cristatella), 1 or 2 upper dorsal scale rows directed upward (vs. 5–7 rows in B. hayeki and 4–10 in B. cristatella), well-developed nuchal crest (weakly-developed in B. cristatella) with cres- cent-shaped scales longer than ED (vs. lanceolate and shorter than ED in B. cristatella), enlarged scales on temporal region (vs. absent in B. hayeki), 3rd finger shorter than fourth (vs. longer in B. hayeki), the orbital area and the tym- panum mostly pale (vs. mostly black in B. hayeki), and tail colouration banded (vs. uniform in B. hayeki).
In addition, Bronchocela jubata is distinguished from other congeners by having the following combination of characters: adults reach a maximum SVL of 141.0 mm in males and 142.0 in females, 9–11 supralabials, 10–12 nuchal crest scales to the level of axilla, 56–73 ventrals, 30–37 lamellae on fourth toe, third finger shorter than the fourth; large lateral scales directed downward anteriorly and straight backwards posteriorly, 1 or 2 upper dorsal scale rows on the lateral body directed upward along the body, mid gular scales enlarged, abdominal scales acumi- nated and enlarged compared to pectoral, keeled temporal scales with some enlarged scales and 6 or 7 rows between orbit and tympanum, tympanum more than half the size of orbit. (Amarasinghe et al. 2022) 
CommentDistribution: Records from Islands Salibabu (Sangihe) Karakelong and Talaud, south east of Mindanao cited by De Roij (1915: 123) could not been verified, but may be correct. A series of 14 specimens from Menado, Sulawesi, in RMNH formerly under B. jubata are redetermined as B. celebensis. B. jubata has been incorrectely reported from the Nicobar Islands (de Roij 1915, Smith 1935) but no voucher specimens exist according to DAS 1999 and own information.The occurrence in Southern India at Pondichery is questionably and may be due to introduced specimens. The species did not occur on Andaman Island (Das 1999). Specimens from Cambodia labelled B. cristatella” are B. jubata [Hallermann 2005]. See map (for Sumatra and N Java) in Amarasinghe et al. 2022: 411 (Fig. 1) 
References
  • Amarasinghe AAT, Kamsi M, Riyanto A, Putra CA, Hallermann J, Andayani N, Abinawanto A, Supriatna J. 2022. Taxonomy, distribution, and conservation status of a rare arboreal lizard, Bronchocela hayeki (Mller, 1928) (Reptilia: Agamidae) from northern Sumatra, Indonesia. Zootaxa 5120 (3):409-422 - get paper here
  • AMARASINGHE, ; A.A. THASUN; IVAN INEICH, AWAL RIYANTO, JAKOB HALLERMANN, NOVIAR ANDAYANI, A. ABINAWANTO, JATNA SUPRIATNA 2022. Taxonomy and distribution of a common arboreal lizard, Bronchocela jubata Duméril & Bibron, 1837 (Reptilia: Agamidae), with designation of its lectotype from Java, Indonesia. Zootaxa 5150 (1): 065–082 - get paper here
  • Berthold,A.A. 1840. Ueber verschiedene neue oder seltene Amphibienarten. Abh. Ges. Wiss. Göttingen 1: 47-72 [1842 ?]
  • Berthold,A.A. 1846. Mittheilungen über das zoologische Museum zu Göttingen. I. Verzeichnisss der aufgestellten Reptilien. Nachr. G. A. Univ. K. Gesell. Wiss. Göttingen 8-10: 1-28
  • Böhme, Wolfgang 2012. About the Collection of and the Research on Agamid Lizards at the Zoologisches Forschungsmuseum Alexander Koenig (ZFMK) in Bonn, Germany. Russ. J. Herpetol. 19 (2): 101-107 - get paper here
  • Böhme, Wolfgang 2014. Herpetology in Bonn. Mertensiella 21. vi + 256 pp. - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Boulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp. - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. 1993. Jour. Sarawak Mus., 44 (65): 128
  • Das, I. 1999. Biogeography of the amphibians and reptiles of the Andaman and Nicobar Islands, India. In: Ota,H. (ed) Tropical Island herpetofauna..., Elsevier, pp. 43-77
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
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  • GRISMER, L. LEE; P.L. WOOD, JR., CHEOL HAENG LEE, EVAN S. H. QUAH, SHAHRUL ANUAR, EHWAN NGADI & JACK W. SITES, JR. 2015. An integrative taxonomic review of the agamid genus Bronchocela (Kuhl, 1820) from Peninsular Malaysia with descriptions of new montane and insular endemics. Zootaxa 3948 (1): 001–023 - get paper here
  • Hallermann, J. 2005. A taxonomic review of the genus Bronchocela (Squamata: Agamidae) with description of a new species from Vietnam. Russ. J. Herpetol. 12 (3): 167-182 - get paper here
  • Iskandar, Djoko T. and Walter R. Erdelen 2006. Conservation of amphibians and reptiles in Indonesia: issues and problems. Amphibian & Reptile Conservation 4 (1): 60-87 - get paper here
  • Janiawati, Ida Ayu Ari; Mirza Dikari Kusrini, Ani Mardiastuti 2016. Structure and Composition of Reptile Communities in Human Modified Landscape in Gianyar Regency, Bali. HAYATI Journal of Biosciences, doi:10.1016/j.hjb.2016.06.006 - get paper here
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  • Manthey U 2008. Agamid lizards of Southern Asia, Draconinae 1. Terralog 7, 160 pp.
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  • MEGANTARA, E. N., JAUHAN, J., SHANIDA, S. S., HUSODO, T., FAUZI, D. A., HENDRAWAN, R., WULANDARI, I. & YUANSAH, Y. 2022. Herpetofauna distribution in different land cover types of West Java, Indonesia. Biodiversitas Journal of Biological Diversity, 23(6) - get paper here
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