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Cachryx defensor (COPE, 1866)

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Higher TaxaIguanidae, Iguania, Sauria, Squamata (lizards) 
Subspecies 
Common NamesYucatán Spinytail Iguana 
SynonymCachryx defensor COPE 1866: 124
Cachryx defensor — BOULENGER 1885: 199
Ctenosaura erythromelas BOULENGER 1886
Cachryx defensor — COPE 1886: 270
Cachryx erythromelas — COPE 1887
Ctenosaura defensor — GÜNTHER 1890
Ctenosaura (Cachryx) annectens WERNER 1911
Ctenosaura erythromelas — BAILEY 1928: 46
Ctenosaura defensor — BAILEY 1928: 48
Enyaliosaurus erythromelas — SMITH & TAYLOR 1950
Enyaliosaurus defensor — DUELLMAN 1965
Ctenosaura defensor — LINER 1994
Ctenosaura defensor — KÖHLER 2000: 75
Ctenosaura (Enyaliosaurus) defensor — KÖHLER et al. 2000
Cachryx defensor — MALONE et al. 2017 
DistributionS Mexico (Campeche, Yucatan, Quintana Roo), Guatemala (Rio Azul)

Type locality: Yucatan. Restricted to Chichén Itzá, Yucatan, Mexico by SMITH & TAYLOR 1950: 77.  
Reproductionoviparous 
TypesSyntypes: USNM 12282
Holotype: BMNH [erythromelas] 
DiagnosisDiagnosis (genus). The following characters differentiate Cachryx, with respect to Ctenosaura: Teeth with 5 or more cusps. Modal number of premaxillary teeth, 5. Crista cranii arranged stepwise between frontal and prefrontal. Subdigital lamellae under the first phalanx of the third digit of the pes not fused. Interparietal (scale covering parietal eye) indistinct or absent. Intercalary scales absent or reduced. Conspicuously enlarged, spiniferous scales present on the anterodorsal surface of the femoral and tibial portions of the hind limbs [from Malone et al. 2017].

Diagnosis (defensor): BAILEY 1928: 48

Diagnosis (erythromelas): BAILEY 1928: 46 
CommentType species: Cachryx defensor COPE 1866: 124 is the type species of the genus Cachryx COPE 1866.

Distribution: the record(s) from Guatemala were originally thought to be C. alfredschmidti but then identified as C. defensor (Malone et al. 2017). 
EtymologyFrom the Latin Cachryo, meaning catkin or parched barley, in reference to the shape of the tail. 
References
  • Alberts, A.C.; Carter, R.L.; Hayes, W.K. & Martins, E.P. (eds.) 2004. Iguanas - Biology and Conservation. University of California Press, 356 pp.
  • Bailey, J. W. 1928. A revision of the lizards of the genus Ctenosaura. Proc. US Natl. Mus. 73 (2733): 1-55. - get paper here
  • Barbour, T., and L. J. Cole. 1906. Vertebrata from Yucatan. Reptilia, Amphibia and Pisces. Bull. Mus. Comp. Zool. Harvard 50: 146-159 - get paper here
  • Boulenger, G. A. 1886. Description of a new iguanoid lizard living in the Society's Gardens. Proc. Zool. Soc. London 1886: 241 - get paper here
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Natural History). Vol. 2, Second edition. London, xiii+497 pp. - get paper here
  • Buckley, Larry J.; Kevin de Queiroz, Tandora D. Grant, Bradford D. Hollingsworth, John B. Iverson (Chair, <br />Stesha A. Pasachnik, and Catherine L. Stephen (Iguana Taxonomy Working Group, ITWG 2016. A Checklist of the Iguanas of the World (Iguanidae; Iguaninae). Herp. Cons. Biol. 11 (Monograph 6) - get paper here
  • Calderon-Mandujano, R. & Mora-Tembre, L. 2004. New distributional records and comments on amphibians and reptiles from Quintana Roo, Mexico. Herpetological Review 35 (3): 295-296 - get paper here
  • Cope, E.D. 1866. Fourth contribution lo the herpetology of tropical America. Proc. Acad. nat. Sci. Philadelphia, 18: 123-132 - get paper here
  • Cope, E.D. 1886. On the species of Iguaninae. Proc. Amer. Philos. Soc. 23 (122): 261-271 - get paper here
  • Duellman,W.E. 1965. Amphibians and reptiles from the Yucatán Peninsula, México. Univ. Kansas Publ. Mus. Nat. Hist. 15: 577-614. - get paper here
  • Köhler, G. 1996. Freilanduntersuchungen zur Morphologie, Verbreitung und Lebensweise des Yucatán-Schwarzleguans (Ctenosaura defensor). Salamandra 32 (3): 153-162 - get paper here
  • Köhler, G. 2000. Reptilien und Amphibien Mittelamerikas, Bd 1: Krokodile, Schildkröten, Echsen. Herpeton Verlag, Offenbach, 158 pp.
  • Köhler, G. 2002. Das Leguanportrait: Ctenosaura defensor COPE 1866. Iguana Rundschreiben 15 (2): 6-8
  • Köhler, G. 2008. Reptiles of Central America. 2nd Ed. Herpeton-Verlag, 400 pp.
  • Köhler, G.; W. Schroth & B. Streit 2000. Systematics of the Ctenosaura group of lizards (Reptilia: Sauria: Iguanidae). Amphibia-Reptilia 21 (2): 177-191 - get paper here
  • Lee, J. C. 2000. A field guide to the amphibians and reptiles of the Maya world. Cornell University Press, Ithaca,
  • Lee, J.C. 1996. The amphibians and reptiles of the Yucatán Peninsula. Comstock, Cornell University Press, Ithaca, 500 pp.
  • Malone, Catherine L.; Víctor Hugo Reynoso, Larry Buckley 2017. Never judge an iguana by its spines: Systematics of the Yucatan spiny tailed iguana, Ctenosaura defensor (Cope, 1866). Molecular Phylogenetics and Evolution 115: 27-39 - get paper here
  • Norell, Mark & De Queiroz, Kevin 1991. The earliest iguanine lizard (Reptilia, Squamata) and its bearing on iguanine phylogeny. American Museum novitates (2997) - get paper here
  • Smith,H.M. & Taylor,E.H. 1950. An annotated checklist and key to the reptiles of Mexico exclusive of the snakes. Bull. US Natl. Mus. 199: 1-253 - get paper here
 
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