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Calotes manamendrai AMARASINGHE & KARUNARATHNA, 2014

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common NamesEnglish: Manamendra-Arachchi’s Whistling Lizard
Sinhala: Manamendra-Arachchigë Uruwan Katussa. 
SynonymCalotes manamendrai AMARASINGHE & KARUNARATHNA in AMARASINGHE et al. 2014 
DistributionC Sri Lanka

Type locality: Riverstone, Knuckles, Sri Lanka; 1000 m elevation  
Reproductionoviparous (by implication) 
TypesHolotype: NMSL5976, adult female, SVL 69.7 mm (collector and the date unknown). Paratype: NMSL5977, adult female, SVL 59.8 mm; other details are the same as the holotype. 
CommentDiagnosis. Females of Calotes manamen- drai sp. nov. are different from the females of Calotes liolepis by having nonenlarged pectoral scales (vs. enlarged); dorsal scales on the body, carinate (vs. smooth); midgular scales smaller than rest of the throat scales (vs. midgular scales same size with throat scales); seven stripes each side on gular area distinct, brown (vs. bands on gular area faint, brown); shoulder pit, black (vs. cream white to brown); and upper arm with carinate ventral scales (vs. smooth). Females of Calotes manamendrai sp. nov. differ from females of Calotes desilvai by having scales on the ventral surface of thigh, carinate (vs. smooth); midgular scales smaller than rest of the throat scales (vs. midgular scales same size with throat scales); seven stripes each side on gular area distinct, brown (vs. bands on gular area, black); shorter AG (141.6%–157.1% of HL) vs. longer AG (174.7%–183.8% of HL); longer upper arm (54.4%–55.4% of HL) vs. shorter upper arm (48.2%–49.2% of HL); Toe V 31.3%–38.6% of HL (vs. 41.9%–45.9% of HL); temporal region with large carinate scales (vs. without large carinate scales); 90–94 ventrals (vs. 72–77); dorsal scales on the body carinate (vs. smooth). Calotes manamendrai sp. nov. further differs from congeners by the following opposing characters: Calotes nigrilabris: ventral scales larger than dorsal; Calotes ceylonensis: lateral scales pointing straight backward; Calotes liocephalus and Calotes pethiyagodai: spines above the tympanum absent; Calotes calotes and Calotes versicolor: lateral scales pointing backward and upward. 
EtymologyThe species epithet is an eponym Latinized in the genitive singular, honoring Kelum Nalinda Manamendra– Arachchi for his generous teaching and guidance in taxonomic studies for the first two authors as well as his remarkable contributions to herpetology, zooarchaeology, and biodiversity conservation in Sri Lanka. 
References
  • Amarasinghe, A.A. Thasun; D.M.S. Suranjan Karunarathna, and J. Fujinuma 2014. A New Calotes Species from Sri Lanka with a Redescription of Calotes liolepis Boulenger, 1885. Herpetologica 70 (3): 323-338 - get paper here
  • Kwet, Axel 2015. Liste der im Jahr 2014 neu beschriebenen Reptilien. Terraria Elaphe 2015 (3): 50-64 - get paper here
 
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