Chelonoidis denticulata (LINNAEUS, 1766)

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Higher Taxa	Testudinidae, Testudines (turtles)
Subspecies
Common Names	E: Yellow-footed Tortoise G: Waldschildkröte
Synonym	Testudo denticulata LINNAEUS 1766 Testudo tabulata WALBAUM 1782 (nomen illegitimum) Testudo tabulata SCHOEPFF 1792 Testudo tessellata SCHNEIDER 1792 Testudo hercules SPIX 1824 Testudo sculpta SPIX 1824 Testudo cagado SPIX 1824 Testudo planata GMELIN 1831 (nomen nudum) Testudo foveolata SCHINZ 1833 (nomen nudum) ? Testudo tabulata — DUMÉRIL & BIBRON 1835: 89 (?) Testudo Tabulata — DUMÉRIL & BIBRON 1854: 220 Geochelone denticulata — WILLIAMS 1960 Geochelone denticulata — DUELLMAN 1978: 194 Chelonoidis denticulata — OBST 1985 Geochelone denticulata — GORZULA & SEÑARIS 1999 Chelonoidis denticulata — MCCORD & JOSEPH-OUNI 2004 Geochelone denticulata — MORAVEC & APARICIO 2005 Chelonoidis denticulata — GERLACH (pers. comm.) Chelonoidis denticulata — LE et al. 2006 Geochelone denticulata — WHITHWORTH & BEIRNE 2011 Chelonoidis denticulata — MCNISH 2011
Distribution	SE Venezuela, Caribbean lowlands of Guyana, French Guiana, and Surinam; Brazil (Amazon Basin, isolated range in E Brazil, Espírito Santo, Pará), E Ecuador, Colombia, NE Peru (Pasco), N/E Bolivia, Trinidad, Guadeloupe. Type locality: in error (see comment) Map legend: - Region according to the TDWG standard, not a precise distribution map. NOTE: TDWG regions are generated automatically from the text in the distribution field and this does not always work properly. We are working on it.
Types	Holotype: NRM (NHRM) DeGeer collection 21, according to Andersson 1900.
Comment	Terra typica: "Virginia," U.S.A.; in error. LOFTIN (1965) reports this species from Panama but assumes that it most likely was an escapee from a pet shop. Reaches 70 cm in carapace length (males), although usually most specimens do not grow beyond 50 cm. Males are smaller than females. Reproduction: oviparous.
References	Andersson, L.G. 1900. Catalogue of Linnean type-specimens of Linnaeus's Reptilia in the Royal Museum in Stockholm. Bihang till Konglika Svenska Vetenskaps-Akademiens. Handlingar. Stockholm. (4) 26 (1): 1-29. Barrio-Amorós, César L. & Brewer-Carias, Charles 2008. Herpetological results of the 2002 expedition to Sarisari–ama, a tepui in Venezuelan Guayana, with the description of five new species. Zootaxa 1942: 1-68 - get paper here Bernarde, Paulo Sérgio;, Reginaldo Assêncio Machado & Luiz Carlos Batista Turci 2011. Herpetofauna da área do Igarapé Esperança na Reserva Extrativista Riozinho da Liberdade, Acre – Brasil. Biota Neotrop. 11 (3): 117-144 - get paper here Bonin, F., Devaux, B. & Dupré, A. 2006. Turtles of the World. English translation by P.C.H. Pritchard. Johns Hopkins University Press, 416 pp. Castaño-Mora, O.V., and M. Lugo R. 1981. Estudio comparativo del comportamiento de dos especies de morrocoy: Geochelone carbonaria y Geochelone denticulata y aspectos comparables de su morfología esterna. Cespedesia 10:55-122 Duellman, W. E. 1978. The biology of an equatorial herpetofauna in Amazonian Ecuador. Misc. Publ. Univ. Kans. Mus. Nat. Hist. 65: 1-352 - get paper here Duellman, W.E., & Salas, A.W. 1991. Annotated checklist of the amphibians and reptiles of Cuzco Amazonico, Peru Occas. Papers Mus. of Natur. Hist., Univ. of Kansas, Lawrence (143): 13 pp. - get paper here Duméril, A. M.C., G. BIBRON & A. DUMÉRIL 1854. Erpétologie générale ou Histoire Naturelle complète des Reptiles. Vol. 9. Paris, XX + 440 S. - get paper here Duméril, A.M.C., and G. Bibron. 1835. Erpétologie Générale ou Histoire Naturelle Complète des Reptiles, Vol. 2. Librairie Encyclopédique de Roret, Paris, iv + 680 p. - get paper here Ernst,C.H. and Barbour,R.W. 1989. Turtles of the World. Smithsonian Institution Press, Washington D.C. - London Ferronato, B. et al. 2011. New locality records for chelonians (Testudines: Chelidae, Podocnemididae, Testudinidae) from Departamento de Pasco, Peru. Herpetology Notes 4: 219-224 Gaffney, E. S. 1979. Comparative cranial morphology of Recent and fossil turtles. Bull. Amer. Mus. Nat. Hist. 164 (2): 67-376 Gorzula Stefan & Senaris J. Celsa 1999. In: Contribution to the herpetofauna of the Venezuelan Guayana. I: a data base. Scientia Guaianae, Caracas, No. 8 [1998], 269+ pp.; ISBN 980-6020-48-0 Gray,J. E. 1866. Notes on some young specimens of tortoises (Testudo). Proc. Zool. Soc. London 1866: 305-306 - get paper here Hoogmoed,M.S. and Gruber,U. 1983. Spix and Wagler type specimens of reptiles and amphibians in the Natural History Musea in Munich (Germany) and Leiden (The Netherlands). Spixiana Suppl. 9: 319-415 - get paper here Knaack, J. 2006. Beobachtungen zum Vorkommen von Geochelone (Chelonoidis) im südlichen Bolivien. Reptilia (Münster) 11 (59): 77-82 - get paper here Le, Minh; Christopher J. Raxworthy, William P. McCord and Lisa Mertz 2006. A molecular phylogeny of tortoises (Testudines: Testudinidae) based on mitochondrial and nuclear genes. Molecular Phylogenetics and Evolution 450 (2): 517-531 - get paper here Lehr, E. 2001. New records for amphibians and reptiles from Departamentos Pasco and Ucayali, Peru. Herpetological Review 32 (2): 130-132 Lehr, E. 2002. Amphibien und Reptilien in Peru. Natur und Tier-Verlag (Münster), 208 pp. - get paper here Linné, C. von [= Linnaeus, C.] 1766. Systema naturæ per regna tria naturæ, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Tomus I. Editio duodecima, reformata. Laurentii Salvii, Stockholm, Holmiae. 1-532 pp. Loftin, H. 1965. On unusual distributional records from the Panama Canal Zone region. Carib. J. Sci. 5 (1-2): 83-86 McCord, W.P. & Joseph-Ouni, M. 2004. Chelonian Illustrations #14: North and South American Tortoises. Reptilia (GB) (34): 47-50 - get paper here McNish, T. 2011. La fauna del archipiélago de San Andrés, Providencia y Santa Catalina, Colombia, Sudamérica. Colomba Andina de Impresos, ISBN 978-958-99518-1-1 Mendes-Pinto, T. J. & S. Marques de Souza 2011. Preliminary assessment of amphibians and reptiles from Floresta Nacional do Trairão, with a new snake record for the Pará state, Brazilian Amazon. Salamandra 47 (4): 199-206 - get paper here Moravec J., Aparicio J. 2005. Notes on the herpetofauna of Bioceanica and Bolpebra (Provincia Nicolas Suaréz, Departamento Pando, Biolivia). Journal of the National Museum (Prague), Natural History Series 174 (1-4): 95-113 Powell, R. & Henderson, R.W. 2003. A second set of addenda to the checklist of West Indian Amphibians ad Reptiles. Herpetological Review 34 (4): 341-345 Pritchard, P.C.H., and P. Trebbau. 1984. The Turtles of Venezuela. SSAR Contributions to Herpetology 2: viii + 402 pp. RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & 2012. Reptiles of Venezuela: an updated and commented checklist Zootaxa 3211: 1–64 - get paper here Schneider, J.G. 1792. Beschreibung und Abbildung einer neuen Art von Wasserschildkröte nebst Bestimmungen einiger bisher wenig bekannten fremden Arten. Schriften der Gesellschaft Naturforschender Freunde zu Berlin 10: 259-283 Silva-Soares, T.; R.B. Ferreira; R.O.L. Salles; C.F.D. Rocha. 2011. Continental, insular and coastal marine reptiles from the municipality of Vitória, state of Espírito Santo, southeastern Brazil. Check List 7 (3): 290-298 - get paper here Stevenson, Pablo R.; Borda, Carlos A.; Rojas, Ana M.; Álvarez, M. 2007. Population size, habitat choice and sexual dimorphism of the Amazonian tortoise (Geochelone denticulata) in Tinigua National Park, Colombia. Amphibia-Reptilia 28 (2): 217-226 - get paper here Valverde, J. 2004. Giant tortoises. Reptilia (GB) (37): 11-19 - get paper here Valverde, J. 2009. Südamerikanische Schildkröten. Reptilia (Münster) 14 (6): 16-27 - get paper here Vargas-Ramírez, Mario Jérôme Maran and Uwe Fritz 2010. Red- and yellow-footed tortoises, Chelonoidis carbonaria and C. denticulata (Reptilia: Testudines: Testudinidae), in South American savannahs and forests: do their phylogeographies reflect distinct habitats? Organisms Diversity & Evolution 10 (1): 161-172 - get paper here Whithworth, A. & Beirne, C. 2011. Reptiles of the Yachana Reserve. Global Vision International, 130 pp. - get paper here
External links	IUCN Red List - Chelonoidis denticulata - Vulnerable, VU http://home.earthlink.net/~rednine/gallery1.htm http://www.omnh.ou.edu/personnel/herpetology/vitt/Herps.html http://www.tortoise.org/archives/carbdent.html Google images

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