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Chironius maculoventris DIXON, WIEST & CEI, 1993

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Higher TaxaColubridae, Colubrinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesCentral Sipo
sexcarinatus: Wagler's Puffing Snake 
SynonymChironius quadricarinatus maculoventris DIXON, WIEST & CEI 1993
Chironius bicarinatus — PETERS & OREJAS-MIRANDA 1970 (part.)
Chironius quadricarinatus maculoventris DIXON, WIEST & CEI 1993: 181
Chironius maculoventris — HOLLIS 2006
Chironius quadricarinatus maculoventris — SCROCCHI et al. 2006: 78
Chironius maculoventris — WALLACH et al. 2014: 161
Chironius quadricarinatus maculoventris — CANO et al. 2015
Chironius maculoventris — TORRES-CARVAJAL et al. 2018 
DistributionNE Argentina (Corrientes, Chaco, Formosa, Santa Fe), W Paraguay (Chaco forests and associated vegetation), Bolivia (Tarija; elevation 440 m, EMBERT 2002), Brazil (Rio Grande do Sul)

Type locality: Corrientes province, Argentina.  
TypesHolotype: ZMUC (was UZMK) 60816 
DiagnosisDiagnosis: brown speckling on yellow·olive ventrals and a bold or darkened postocular stripe versus immaculale yellow ventrals and no noticeable postocular stripe in the nominate form [DIXON, WIEST & CEI 1993: 182]

Diagnosis (quadricarinatus): This subspecies of C. quadricarinatus has uniform yellow ventrals and subcaudals and no bold postocular slripe, versus brown speckling on yellow-olive ventrals and a distinct dark postocular stripe in C q. maculoventris [DIXON, WIEST & CEI 1993: 185] 
CommentSynonymy that of PETERS & OREJAS-MIRANDA 1970 and CEI 1993. GIRAUDO & SCROCCHI 2002, following DIXON et al. 1993, state that old material identified as Herpetodryas sexcarinatus is actually Chironius quadricarinatus maculoventris and thus this species does not occur in Argentina. 
  • Cacciali, Pier; Hugo Cabral 2015. The genus Chironius (Serpentes, Colubridae) in Paraguay: composition, distribution, and morphology. Basic and Applied Herpetology 29: - get paper here
  • Cacciali, Pier; Norman J. Scott, Aida Luz Aquino Ortíz, Lee A. Fitzgerald, and Paul Smith 2016. The Reptiles of Paraguay: Literature, Distribution, and an Annotated Taxonomic Checklist SPECIAL PUBLICATION OF THE MUSEUM OF SOUTHWESTERN BIOLOGY, NUMBER 11: 1–373 - get paper here
  • Cano, Paula Daniela; Héctor Alejandro Ball, Matías Federico Carpinetto, Germán Darío Peña 2015. Reptile checklist of Río Pilcomayo National Park, Formosa, Argentina. Check List 11 (3): 1658 - get paper here
  • Dixon J R; Wiest J A Jr; Cei J M 1993. Revision of the Neotropical snake genus Chironius Fitzinger (Serpentes, Colubridae). MUSEO REGIONALE DI SCIENZE NATURALI MONOGRAFIE (TURIN) 13 :1-280
  • Embert, Dirk 2002. Geographic distribution. Chironius quadricarinatus maculoventris (Cobra Cipo, Sacaiboia). Herpetological Review 33 (3): 226-227 - get paper here
  • Hollis, J.L. 2006. Phylogenetics of the genus Chironius Fitzinger 1826 (Serpenes, Colubridae) based on morphology. Herpetologica 62 (4): 435-452 - get paper here
  • Marra Santos, Fidélis Júnio; Márcio Repenning, Christian Beier, Glaucia Maria Funk Pontes 2015. First record of Chironius maculoventris Dixon, Wiest & Cei, 1993 (Squamata: Serpentes: Colubridae) in Brazil PDF Herpetology Notes 8: 169-171 - get paper here
  • Pesci, Giuliano; Sanchez, José Manuel; Muniz Leão, Suelem; Pelegrin, Nicolás 2018. Reptiles y anfibios de una localidad del Chaco Húmedo en Formosa, Argentina. Cuad. herpetol. 32 (1): 47-54 - get paper here
  • Peters, James A.; Donoso-Barros, Roberto & Orejas-Miranda, Braulio 1970. Catalogue of the Neotropical Squamata: Part I Snakes. Part II Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 347 pp. - get paper here
  • Scrocchi, G.J.; Moreta, J.C. & Kretzschmar, S. 2006. Serpientes del Noroeste Argento [Jujuy, Salta, Tucumán, Catamarca, La Rioja, Santiago del Estero]. Fundación Miguel Lillo, Tucumán, 178 pp.
  • Torres-Carvajal, O., Echevarría, L.Y., Lobos, S.E., Venegas, P.J., Kok, P.J.R. 2018. Phylogeny, diversity and biogeography of Neotropical sipo snakes (Serpentes: Colubrinae: Chironius). Molecular Phylogenetics and Evolution 130: 315-329 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
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