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Clelia equatoriana (AMARAL, 1924)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Common NamesE: Equatorial Mussurana 
SynonymBarbourina equatoriana AMARAL 1924: 201
Clelia clelia scytalina — DUNN 1944: 201
Clelia equatoriana — PETERS & OREJAS-MIRANDA 1970: 64
Clelia equatoriana — ZAHER 1996
Clelia equatoriana — WALLACH et al. 2014: 170 
DistributionE Costa Rica, E Panama, NW Ecuador, Colombia (Caldas,Valle del Cauca etc.), Peru (Piura)

Type locality: Guayaquil, Ecuador  
TypesHolotype: USNM 62790, male 
DiagnosisOriginal description: “Snout round and scarcely projecting. Rostral a little deeper than broad, its portion visible from above equal to 1/3 its distance from the frontal; internasals twice as broad as long, half as long as the praefrontals; frontal a little longer than broad and longer than its distance from the end of the snout, a little shorter than the parietals; nasal entire, in contact with the praeocular (on the right) which does not reach the upper surface of the head; no loreal; 2 small postoculars, subequal; temporals 2 + 3; 7 upper labials, the 3d and 4th entering the orbit; 2d touching the prefrontal on the left; 1st lower labials in contact with its fellow behind the symphysial; 4 lower labials in contact with the anterior chin-shields which are a little longer than the posterior. Scales in 17 rows. Ventrals 204, rounded laterally; anal entire; subcaudals 75 pairs.” (Amaral 1924)

Coloration: “Pinkish-yellow (red?) above, all the scales tipped with black; head brown above, with a yellowish-white band occupying the parietals, temporals, the 3 posterior upper labials, and the occiput, and followed by a dark blotch on the nape; chin dark; throat and lower surface of the body and tail yellowish-white.” (Amaral 1924) 
CommentSimilar species: Clelia equatoriana is often confounded with Clelia clelia. However, Clelia equatoriana has dorsal scale in 17 rows at midbody vs 19 in Clelia clelia (Peters & Orejas-Miranda 1970).

Type species: Barbourina equatoriana AMARAL 1924: 201 is the type species of the genus Barbourina AMARAL 1924, now a synonym of Clelia. 
  • Amaral, A. do 1924. New genus and species of South American snakes contained in the United States National Museum. J. Washington Acad. Sci. 14: 200-202 - get paper here
  • Arteaga AF, Bustamante-Enríquez LM and Guayasamin JM 2013. The Amphibians and Reptiles of Mindo. - get paper here
  • Castro-Herrera, F. & Vargas-Salinas, F. 2008. Anfibios y reptiles en el departamento del Valle del Cauca, Colombia. Biota Colombiana 9 (2): 251 - 277 - get paper here
  • Chávez-Arribasplata, Juan C.; Diego Vásquez, Claudia Torres, Lourdes Y. Echevarría, and Pablo J. Venegas 2016. Confirming the presence of Clelia equatoriana Amaral, 1924 (Squamata: Dipsadidae) in Peru. Amphibian & Reptile Conservation 10 (1): 1–4 - get paper here
  • Dunn,E.R. 1944. Los géneros de anfibios y reptiles de Colombia, III. Orden de las serpientes. Caldasia 3 (12): 155-224 - get paper here
  • Köhler, G. 2001. Reptilien und Amphibien Mittelamerikas, Bd. 2: Schlangen. Herpeton-Verlag (Offenbach), 174 pp.
  • Morato, S.A.A.; Franco, F.L. & Sanches, E.J. 2003. Uma nova espécie de Clelia (Serpentes, Colubridae) do sul do Brasil. Phyllomedusa 2 (2): 93-100 - get paper here
  • Pérez-santos, C. E. 1986. Zoogeografía de los Ophidia en Colombia. Univ. Complutense, Madrid, 2 vols., 971 pp.
  • Pérez-Santos,C. & Moreno, A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Rojas-Morales, Julián Andrés 2013. Description of ophiophagy in Clelia equatoriana (Amaral, 1924) (Serpentes: Dipsadidae) in captivity. Herpetology Notes 6: 425-426.
  • Rojas-Morales, Julián Andrés, Héctor Fabio Arias-Monsalve y Gustavo A. González-Durán 2014. Anibios y reptiles de la región centro-sur del departamento de Caldas, Colombia. Biota Colombiana 15 (1): 73-93 - get paper here
  • Rojas-Morales, Julián Andrés 2012. Snakes of an urban-rural landscape in the central Andes of Colombia: species composition, distribution, and natural history. Phyllomedusa 11 (2): 135-
  • Torres-Carvajal O, Pazmiño-Otamendi G, Salazar-Valenzuela D. 2019. Reptiles of Ecuador: a resource-rich portal, with a dynamic checklist and photographic guides. Amphibian & Reptile Conservation 13 (1): [General Section]: 209–229 (e178) - get paper here
  • Valencia-Zuleta A, Jaramillo-Martínez AF, Echeverry-Bocanegra A, Viáfara-Vega R, Hernández-Córdoba O, Cardona-Botero VE, Gutiérrez-Zúñiga J, Castro-Herrera F. 2014. Conservation status of the herpetofauna, protected areas, and current problems in Valle del Cauca, Colombia. Amphibian & Reptile Conservation 8 (2): 1–18 (e87) - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zaher, H. 1996. A new genus and species of pseudoboine snake, with a revision of the genus Clelia (Serpentes, Xenodontinae). Boll. Mus. Reg. Sci. Nat. Torino 14: 289-337.
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