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Cnemaspis kohukumburai KARUNARATHNA, POYARKOV, DE SILVA, MADAWALA, BOTEJUE, GORIN, SURASINGHE, GABADAGE, UKUWELA & BAUER, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Kohukumbures’ day gecko
Sinhala: Kohukumburege diva-seri hoona 
SynonymCnemaspis kohukumburai KARUNARATHNA, POYARKOV, DE SILVA, MADAWALA, BOTEJUE, GORIN, SURASINGHE, GABADAGE, UKUWELA & BAUER 2019 
DistributionSri Lanka (Kandy District)

Type locality: granite tunnel in Kadugannawa, Kandy District,
Central Province, Sri Lanka (7.251800° N, 80.509378° E,
WGS1984; elevation 427 m.  
Reproduction 
TypesHolotype. NMSL.2019.05.01, adult male, 33.9 mm SVL (Fig. 10),
collected around 09.00 hrs) on 12 October 2018 by Suranjan Karunarathna and Anslem de Silva.
Paratypes. NMSL.2019.05.02, adult male, 32.5 mm SVL, and
NMSL.2019.05.03, adult female, 31.0 mm SVL collected from a granite tunnel in Kadugannawa, Kandy District, Central Province,
Sri Lanka (7.251658° N, 80.512561° E, WGS1984; elevation
435 m; around 10.00 hrs), on 12 October 2018 by Suranjan
Karunarathna and Anslem de Silva. 
DiagnosisDiagnosis. Cnemaspis kohukumburai sp. nov. can be readily distinguished from its Sri Lankan congeners by a combination of the following morphological and meristic characteristics and color pattern: maximum SVL 33.9 mm; dorsum with homogeneous, smooth granular scales; 2/2 supranasals, 1 internasal and 1/1 postnasal present; 3 enlarged postmentals; postmentals bounded by 5 chin scales; chin and gular scales smooth, granular, juxtaposed; pectoral and abdominal scales smooth, subimbricate; 7 – 8 well developed tubercles on posterior flank; 150 – 159 paravertebral granules linearly arranged; 23 belly scales across the venter; precloacal pores absent in males, 6 – 9 femoral pores on each side in males separated by 25 unpored femoro-precloacal scales, 1 – 2 unpored posterior femoral scales; 131 – 134 ventral scales; 81 – 88 midbody scales; subcaudals smooth, subhexagonal, enlarged, subequal, forming aregular median row; 8 – 9 supralabials; 7 – 8 infralabials; 21 – 22 total lamellae on digit IV of manus, and 23 – 25 total lamellae on digit IV of pes.

Comparisons. The new species differs from species of the C. podihuna clade sensu Agarwal et al. (2017) as follows: from C. kandambyi, C. molligodai and C. podihuna by the absence (versus presence) of precloacal pores; from C. gemunu, C. godagedarai, C. phillipsi, C. rammalensis and C. scalpensis by the presence of fewer femoral pores (6 – 9 versus 11 – 14, 12 – 13, 15 – 16, 15 and 13 – 15, respectively); it further differs from C. gemunu and C. scalpensis by the presence of a greater number of belly scales (23 versus 13 – 16 and 17 – 19, respectively), from C. rammalensis by the presence of fewer ventral scales (131 – 134 versus 186 – 207), and from C. phillipsi by the presence of more total lamellae on digit IV of pes (23 – 25 versus 17 – 19). It can be diagnosed from C. alwisi and C. rajakarunai by the presence of fewer ventral scales (131 – 134 versus 145 – 153 and 146 – 186, respectively), and by the presence of more unpored femoro-precloacal scales (25 versus 18 – 19 and 20 – 22, respectively). It can be differentiated from C. punctata by the greater number of paravertebral granules (150 – 159 versus 83 – 91), by the presence of more total lamellae on digit IV of manus (21 – 22 versus 17 – 18), and by fewer well-developed tubercles on the posterior flank (7 – 8 versus 11 – 13). Further, it can be distinguished from C. nilgala by the presence of a greater number of belly scales (23 versus 17 – 19) across the midbody, and by the presence of more total lamellae on digit IV of pes (23 – 25 versus 17 – 19). Cnemaspis kohukumburai sp. nov. also clearly differs by the absence of precloacal pores and the presence of clearly enlarged, hexagonal or subhexagonal subcaudal scales from the following species of the C. kandiana clade sensu Agarwal et al. (2017): C. amith, C. gotaimbarai sp. nov., C. ingerorum, C. kallima, C. kandiana, C. kumarasinghei, C. latha, C. menikay, C. nandimithrai sp. nov., C. pava, C. pulchra, C. retigalensis, C. samanalensis, C. silvula, C. tropidogaster and C. upendrai. 
Comment 
EtymologyThe specific epithet is an eponym Latinized (kohukumburai) in the masculine genitive singular, honouring the Sri Lankan warrior ‘Kohukumbure Walauwe Rate Rala’ for his valiant feats in the Great Rebellion of 1817 – 1818 which was initiated in Uva-Wellassa. 
References
  • Karunarathna, Suranjan; Nikolay A. Poyarkov, Anslem de Silva, Majintha Madawala, Madhava Botejue, Vladislav A. Gorin, Thilina Surasinghe, Dinesh Gabadage, Kanishka D.B. Ukuwela & Aaron M. Bauer 2019. Integrative taxonomy reveals six new species of day geckos of the genus Cnemaspis Strauch, 1887 (Reptilia: Squamata: Gekkonidae) from geographically-isolated hill forests in Sri Lanka. Vertebrate Zoology 69 (3): 247–298 - get paper here
 
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