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Cnemaspis nigridia (SMITH, 1925)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Borneo Black Gecko, Black-spotted Rock Gecko
G: Schwarzflecken-Dünnfingergecko 
SynonymGonatodes nigridius SMITH 1925: 22
Cnemaspis nigridius — BRONGERSMA 1934: 165
Cnemaspis nigridius — KLUGE 1993
Cnemaspis nigridius — MANTHEY & GROSSMANN 1997: 214
Cnemaspis nigridius — COX et al. 1998: 90
Cnemaspis cf. nigridia — CHAN-ARD et al. 1999: 104
Cnemaspis (Cnemaspis) nigridius — RÖSLER 2000: 63
Cnemaspis nigridia — DAS & GRISMER 2003
Cnemaspis nigridia — NAMING & DAS 2004
Cnemaspis nigridius — AULIYA 2006
Cnemaspis nigridia — WOOD et al. 2013 
DistributionIndonesia (Borneo, Sarawak, Mount Gardin)

Type locality: “Mt. Gadin” [Sarawak, Borneo]  
Reproductionoviparous 
TypesHolotype: BMNH 1946.8.22.90 
DiagnosisDiagnosis: Maximum SVL 75.5 mm; 10 or 11 supralabials; 9–11 infralabials; ventral scales weakly to moderately keeled; 10–16 contiguous, pore-bearing, precloacal scales with round pores; 39–43 paravertebral tubercles; tubercles semi-linear to randomly arranged, absent from flanks and lateral caudal furrows; ventrolateral caudal tubercles present; lateral row of caudal tubercles present; caudal tubercles not encircling tail; subcaudals keeled but bearing an enlarged median row of smooth scales; 2–4 postcloacal tubercles on each side of tail base; no enlarged femoral or subtibial scales; submetatarsal scales of first toe enlarged; subtibials keeled; 26–29 subdigital fourth toe lamellae; a pair of large, round, black spots in shoulder region; yellow to white bars on flanks; wide, black and dull-yellow caudal bands (Tables 6,7 in GRISMER et al. 2014). 
CommentDistribution: Reports from Pulau Tioman represent actually C. limi according to GROSSMANN & TILLACK 2004. See map in Grismer et al. 2014: 18 (Fig. 4), Kurita et al. 2017: 526 (Fig. 1). 
References
  • Auliya, M. 2006. Taxonomy, Life History, and conservation of giant reptiles in west Kalimantan. Natur und Tier Verlag, Münster, 432 pp. - get paper here
  • Brongersma,L.D. 1934. Contributions to Indo-Australian herpetology. Zoologische Mededelingen 17: 161-251 - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das I 1994. Cnemaspis gordongekkoi, a new gecko from Lombok, Indonesia, and the biogeography of oriental species of Cnemaspis (Squamata: Sauria: Gekkonidae). Hamadryad 18: 1-9 [1993] - get paper here
  • Das, I. 2005. Revision of the Genus Cnemaspis Strauch, 1887 (Sauria: Gekkonidae), from the Mentawai and Adjacent Archipelagos off Western Sumatra, Indonesia, with the Description of Four New Species. Journal of Herpetology 39 (2): 233–247 - get paper here
  • Das, I. & Grismer, L.L. 2003. Two new species of Cnemaspis Strauch 1887 (Squamata, Gekkonidae) from the Seribuat Archipelago, Pahang and Johor States, West Malaysia. Herpetologica 59 (4): 544-552 - get paper here
  • Das,I. & A. M. BAUER 1998. Systematics and biogeography of Bornean geckos of the genus Cnemaspis Strauch, 1887 (Sauria: Gekkonidae), with the description of a new species. Raffles Bull. Zool. 46 (1): 11-28. - get paper here
  • Grismer, J.L.; Grismer, L.L. & Chav T. 2010. New Species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Southwestern Cambodia. Journal of Herpetology 44 (1): 28–36 - get paper here
  • Grismer, L. Lee & Chan K. Onn 2008. A new species of Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from Pulau Perhentian Besar, Terengganu, Peninsular Malaysia. Zootaxa 1771: 1–15 - get paper here
  • Grismer, L. Lee; Chan K. Onn; Nasir, N. & Sumontha, M. 2008. A new species of karst dwelling gecko (genus Cnemaspis Strauch 1887) from the border region of Thailand and Peninsular Malaysia. Zootaxa 1875: 51–68 - get paper here
  • GRISMER, L. LEE; MONTRI SUMONTHA, MICHAEL COTA, JESSE L. GRISMER, PERRY L. WOOD, JR., OLIVIER S. G. PAUWELS & KIRATI KUNYA 2010. A revision and redescription of the rock gecko Cnemaspis siamensis (Taylor 1925) (Squamata: Gekkonidae) from Peninsular Thailand with descriptions of seven new species. Zootaxa 2576: 1–55 - get paper here
  • Grismer, L.L. & Das, I. 2006. A new species of gekkonid lizard of the genus Cnemaspis STRAUCH 1887 from Pulau Pemanggil, Johor, West Malaysia. Herpetological Natural History 10 (1): 1-7 - get paper here
  • Grismer, L.L. & Ngo, V.T. 2007. Four new species of the gekkonid genus Cnemaspis Strauch 1887 (Reptilia: Squamata) from Southern Vietnam. Herpetologica 63 (4): 482-500 - get paper here
  • Grismer, L.L.; PERRY JR L. WOOD, SHAHRUL ANUAR, AWAL RIYANTO, NORHAYATI AHMAD, MOHD A. MUIN, MONTRI SUMONTHA, JESSE L. GRISMER, CHAN KIN ONN, EVAN S. H. QUAH, OLIVIER S. A. PAUWELS 2014. Systematics and natural history of Southeast Asian Rock Geckos (genus Cnemaspis Strauch, 1887) with descriptions of eight new species from Malaysia, Thailand, and Indonesia. Zootaxa 3880 (1): 001–147 - get paper here
  • Grossmann, W. & Tillack, F. 2004. Pulau Tioman - Perle im Südchinesischen Meer, Teil 1. Reptilia (Münster) 9 (50): 42-49 - get paper here
  • Hien,P. Grossmann,W. & Schäfer, C. 2001. Beitrag zur Kenntnis der landbewohnenden Reptilienfauna von Pulau Tioman, West-Malaysia. Sauria 23 (4): 11-28 - get paper here
  • KURITA, TAKAKI; KANTO NISHIKAWA, MASAFUMI MATSUI, TSUTOMU HIKIDA 2017. A new species of Rock Gecko genus Cnemaspis (Squamata: Gekkonidae) from Western Sarawak, Malaysia. Zootaxa 4258 (6): 525–538 - get paper here
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • Naming, M. & Das, I. 2004. Herpetofauna. In: Sarawak Bau Limestone Biodiversity (H.S. Yong, F.S.P. Ng & E.E. L. Yen, eds.). Sarawak Museum Journal LIX (No. 80; special issue 6): 251-265
  • Nguyen, H.N., Hung, C., Yang, M. et al. 2020. Sympatric competitors have driven the evolution of temporal activity patterns in Cnemaspis geckos in Southeast Asia. Sci Rep 10: 27 - get paper here
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Smith, M. A. 1925. Contribution to the herpetology of Borneo. Sarawak Mus. J. 3 (8): 15-34. - get paper here
  • WOOD, PERRY JR. L.; EVAN S.H. QUAH, SHAHRUL ANUAR M.S., MOHD ABDUL MUIN 2013. A new species of lowland karst dwelling Cnemaspis Strauch 1887 (Squamata: Gekkonidae) from northwestern Peninsular Malaysia. Zootaxa 3691 (5): 538–558 - get paper here
 
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