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Cnemaspis thayawthadangyi LEE, MILLER, ZUG & MULCAHY, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Thayawthadangyi Islands Rock Gecko 
SynonymCnemaspis thayawthadangyi LEE, MILLER, ZUG & MULCAHY 2019 
DistributionMyanmar (Tanintharyi Region)

Type locality: Za Let Aw (Escape Bay), southeast side of Linn Lune Kyun, Thayawthadangyi Kyun Group, Myeik Archipelago, Tanintharyi Region, Myanmar (12.30849°N, 98.0043°E; 80 m elevation).  
Reproduction 
TypesHolotype. USNM 595053 (GenBank accession no. MN104951), adult male collected by Daniel G. Mulcahy and Grant Connette on 12 May 2017, 2100–2230 h.
Paratype. USNM 595052 (GenBank accession no. MN104950), adult male, same collection data as the holotype. 
DiagnosisDiagnosis. Cnemaspis thayawthadangyi sp. nov. is a small diminutive member of the C. kandiana species group and can be distinguished from all other members of the group by a combination of the following morpho- logical characters: 1) a maximum SVL of 29.9 mm; 2) each postmental bordered posteriorly by four scales; 3) four spine-like ventrolateral tubercles on flank; 4) gular scales keeled; 5) pectoral scales and abdominal scales keeled; 6) ventral scales keeled; 7) three precloacal pores; 8) four femoral pores on each leg; 9) subcaudal scales smooth, scales on median row enlarged and smooth; 10) coloration of the gular region silver with dark-gray irregularly shaped streaks; 11) 6–7 supralabials; 12) 6–7 infralabials; 13) 16–18 subdigital lamellae on the 4th toe; 14) 18–20 ventral scales at midbody.

Comparisons. We compare the species Cnemaspis thayawthadangyi sp. nov. to all other members of the Cne- maspis kandiana group from Southern Thailand, Sumatra and islands off its west coast and the Bay of Bengal. Further comparisons can be found in Table 3. C. thayawthadangyi sp. nov. is distinguished from C. phuketensis by having an enlarged medial subcaudal row (versus subcaudals equal in size), precloacal and femoral pores in males (versus both absent), smooth gular scales (versus keeled) and by four scales bounding each postmental scale (versus three). For distinctions between C. tanintharyi sp. nov. and C. thayawthadangyi sp. nov., see the compari- sons section in the species account for C. tanintharyi sp. nov. From the Cnemaspis found in the Bay of Bengal, C. thayawthadangyi sp. nov. is distinguished from C. andersonii by having four bounded scales bordering the post- mental (versus three scales), ventral scales throughout belly keeled (versus all ventral scales smooth), the presence of enlarged medial subcaudal row (versus subcaudals equal sized) and a blotched color pattern (versus immaculate coloration) as well as C. wicksi by three precloacal pores (versus 4–5), 16–18 subdigital lamellae on 4th toe (versus 14), keeled abdominal scales (versus smooth), ventral scales on thigh weakly keeled (versus smooth), and a blotched color pattern (versus immaculate coloration).
The presence of keeled ventral scales on the thigh distinguishes C. thayawthadangyi sp. nov. from all members of the C. kandiana species group found in Sumatra and islands off its west coast (C. aceh, C. andalas, C. phuketen- sis, C. minang, and C. tapanuli) except for C. dezwaani, C. jacobsoni, C. modiglianii, C. pagai and C. whittenorum. Cnemaspis thayawthadangyi sp. nov. is distinguished from C. dezwaani by having three precloacal pores (versus 8–12) and the absence of a pale vertebral stripe (versus present), from C. jacobsoni by having femoral and precloa- cal pores (versus absent), 16–18 subdigital lamellae on 4th toe (versus 14), from C. modiglianii by having 18–20 ventral scales at midbody (versus 24–26); only four precloacal pores (versus 2–4), only four scales surrounding each postmental (versus 3–5) and by a gular coloration consisting of thick dark irregular stripes (versus unpatterned gular coloration), from C. pagai by having 16–18 subdigital lamellae on 4th toe (versus 20), three precloacal pores (versus two), four femoral pores (versus three) and four scales bordering each postmental (versus three), from C. whitteno- rum by having three precloacal pores (versus four), four femoral pores (versus six), 6–7 supralabials (versus 5–6), and the absence of a pale vertebral stripe (versus present). 
CommentHabitat: The two specimens were collected on large tree trunks at night close to a stream. This species is probably more widely distributed than currently documented and likely inhabits similar microhabitats on this and other islands in the group. 
Etymology The specific epithet “thayawthadangyi” is a reference to the type locality of this species—the Thayawthadangyi Kyun (Island) group in the Myeik Archipelago. 
References
  • LEE, JUSTIN L.; ARYEH H. MILLER, GEORGE R. ZUG, DANIEL G. MULCAHY 2019. The discovery of Rock Geckos Cnemaspis Strauch, 1887 (Squamata: Gekkonidae) in the Tanintharyi Region, Myanmar with the description of two new species. Zootaxa 4661 (1): 040–064 - get paper here
 
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