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Crenadactylus rostralis (STORR, 1978)

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Higher TaxaDiplodactylidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesSouth-west Kimberley clawless gecko 
SynonymCrenadactylus ocellatus rostralis STORR 1978
Crenadactylus rostralis - WELLS & WELLINGTON 1984
Crenadactylus ocellatus rostralis — RÖSLER 2000: 64
Crenadactylus rostralis — DOUGHTY et al. 2016 
DistributionAustralia (Western Australia: S Kimberleys)

Type locality: Geikie Gorge, Western Australia (18°07'S, 125°39'E) Map legend:
Type locality - Type locality.
 
Reproductionoviparous 
TypesHolotype. WAM R32154, adult male, collected by W.H. Butler, April 1968. Fixed in 10% formalin, stored in 70% ethanol at WAM.
Paratypes (17). WAM R23058*, 43 km SE Halls Creek, WA (18°27'S, 127°15'E); WAM R26824–26, Napier Range, WA (17°20'S, 124°50'E); WAM R27388–92, Mount Anderson, WA (18°02'S, 123°56'E); WAM R46058– 59*, near Wolfe Creek Meteorite Crater, 122 km SW Halls Creek, WA (18°43'S, 126°45'E); WAM R46115*, 11 km E Margaret River Homestead, WA (18°38'S, 126°58'E); WAM R56841–45*, Lake Argyle, WA (16°15'S, 128°45'E).
Some registration numbers presented in the original description are in error, with WAM R56441–45 associated with different species. The error originates from a minor numerical error in the registration numbers presented by Storr (1978), and underlined here. The correct numbers associated with the paratype specimens are WAM R56841– 45. Nine paratypes (marked with an asterisk) designated by Storr (1978) fall within the distribution of lineages associated with the C. naso complex identified by Oliver et al. (2012b), and are not true C. rostralis (Doughty et al. 2016: 262). 
CommentDiagnosis. A moderate-sized (to 32.7 mm SVL) species of Crenadactylus. Rostral not in contact with nostril, 2–3 internasals present, not extending beyond supranasal, postmentals granular, dorsal scales homogeneous and weakly keeled, 4–6 pre-cloacal pores, innermost pore-bearing scales in contact, no enlarged tubercles on original tails. Ground colour light greyish brown; dorsal pattern consists of well-defined pale and dark longitudinal stripes, lateral zone light grey with usually two moderate to well-defined dark lines between limbs, stippled with single pale scales; ventrum pale grey to white with moderate stippling forming longitudinal lines.

Variation. Other specimens generally corresponded to the holotype specimen, with the following exceptions. The degree of keeling on the dorsal scales ranged from feebly keeled to smooth and rounded. Females lacked pre-cloacal pores and the spurs were smaller and less projecting.
Colouration and pattern. In life (based on recent photographs by PMO), background colour grey-brown with a series of alternating pale and dark alternating longitudinal stripes; pale grey vertebral stripe bordered by medium dark brown paravertebral stripes; pale to medium grey dorsolateral zone, continuing forwards through eye as pale canthal stripe; dark brown lateral stripe, continuing through eye on snout as poorly-defined loreal stripe; light grey ventrolateral zone with occasionally poorly-defined thin dark stripe near ventrolateral edge of body; top of head dark brown with some pale mottling; labials pale, stippled with dark brown. Dark brown paravertebral and lateral stripes usually with scattered pale scales. Lateral stripe usually darker than paravertebral stripes, but can also be of similar shade. Ventro- lateral zone can be much paler than dorsolateral zone. Limbs dark brown with variable small pale spots; original tails continue dorsal pattern, regenerated tails with a mixture of pale and dark scales not forming conspicuous lines.
In preservative, pattern consists of alternating dark and light stripes. Silvery-grey vertebral stripe, narrow, 2–3 scales wide; medium grey-brown paravertebral stripes, bordered by dark brown edges, 6–7 scales wide; pale dorsolateral stripe, 4–5 scales wide, starts at postero-dorsal edge of eye, continues above limbs to tail; dark brown dorso-lateral stripe extends posteriorly from back of eye to top of hindlimb, anteriorly extends to top of ear opening; below dorsolateral line are two thin, ventro-lateral stripes; pale lateral zone divided by thin dark lateral stripe between limbs, forming pale line below dorsolateral that extends from angle of jaw across top of forelimbs and through hindlimb. Ventrum lightly stippled, one or two weakly defined brown lines towards outer edge of ventrum. Pale canthal and dark loreal stripes weakly defined, continuation of pale and dark dorsolateral stripes; top of eyes purplish; labials pale and heavily stippled with brown flecks; posterior edge of jaw heavily stippled, continuing as lateral line below ear opening to forelimb. Limbs dark, back of thigh has strong pale stripe, diffuse with small clusters of pale scales.

Distribution: See map in Doughty et al. 2016: Fig. 2.

Habitat. The few records with collection details all mention occurring in spinifex, including a number of recent samples that were raked from large clumps on and around limestone ranges (Doughty et al. 2016). 
EtymologyThe name rostralis is from the Latin rostrum, meaning beak or snout, in reference to this species’ distinctive configuration of having the rostral scale separated from the nostril. 
References
  • DOUGHTY, PAUL; RYAN J. ELLIS, PAUL M. OLIVER 2016. Many things come in small packages: Revision of the clawless geckos (Crenadactylus: Diplodactylidae) of Australia. Zootaxa 4168 (2): 239–278
  • Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
  • Storr, G.M., & Harold, G. 1978. Herpetofauna of the Shark Bay Region, Western Australia. Rec. West. Aust. Mus. 6 (4): 449-467. - get paper here
  • Wells R W; Wellington C R 1984. A synopsis of the class Reptilia in Australia. Australian Journal of Herpetology 1 (3-4): 73-129 [1983]
 
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