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Cryptoblepharus pulcher (STERNFELD, 1918)

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Higher TaxaScincidae, Eugongylinae, Scincoidea, Sauria, Squamata (lizards) 
SubspeciesCryptoblepharus pulcher clarus (STORR 1961)
Cryptoblepharus pulcher pulcher (STERNFELD 1918) 
Common NamesElegant snake-eyed skink
Bright snake-eyed skink (clarus) 
SynonymAblepharus boutoni pulcher STERNFELD 1918: 423
Ablepharus boutoni virgatus — MERTENS 1931: 112
Ablepharus boutoni virgatus — WORRELL 1963: 35
Cryptoblepharus boutonii virgatus – COOK 1973: 15
Cryptoblepharus virgatus — GREER 1974: 21
Cryptoblepharus virgatus - COGGER 1983: 142
Cryptoblepharus virgatus virgatus – WILSON & KNOWLES 1988: 120
Cryptoblepharus virgatus – STORR et al. 1981: 25
Cryptoblepharus virgatus – COGGER et al. 1983a: 142
Cryptoblepharus virgatus – COVACEVICH & COUPER 1991: 357
Cryptoblepharus virgatus – EHMANN 1992: 183
Cryptoblepharus virgatus – STORR et al. 1999: 25
Cryptoblepharus virgatus – COGGER 2000: 406
Cryptoblepharus virgatus – DALY et al. 2001: 85
Cryptoblepharus virgatus – GREER & JEFFERYS 2001: 3
Cryptoblepharus virgatus – WILSON & SWAN 2003: 150
Cryptoblepharus suburbia WELLS & WELLINGTON, 1985: 27
Cryptoblepharus virgatus suburbia WELLS & WELLINGTON, 1985
Cryptoblepharus virgatus suburbia – WELLS & WELLINGTON 1989: 29
Cryptoblepharus virgatus — COGGER 2000: 406
Cryptoblepharus virgatus — HORNER 2007

Cryptoblepharus pulcher clarus (STORR 1961)
Ablepharus boutonii clarus STORR 1961: 176
Ablepharus boutonii clarus — WORRELL 1963: 35
Cryptoblepharus virgatus clarus – STORR 1976: 55
Cryptoblepharus virgatus clarus – WILSON & KNOWLES 1988: 120.
Cryptoblepharus clarus – WELLS & WELLINGTON 1985: 27
Cryptoblepharus virgatus clarus — STORR et al. 1999
Cryptoblepharus virgatus clarus — WWW 2006
Cryptoblepharus pulcher clarus — HORNER 2007 
DistributionAustralia (far eastern Australia and far southern coastal regions of Western and South Australia, New South Wales).

Type locality: “Neuholland” Cooktown, Qld. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesLectotype: SMF 15680-15681, from Australia, designated by Mertens (1967) (Ablepharus boutoni pulcher)
Syntypes: WAM R18228-31, from lower Dalyup River, in 33° 45’ S, 121° 32’ E (20 mi WNW of Esperance), W. A.; paratype: QM [Ablepharus boutonii clarus] 
CommentSynonymy after HORNER 2007.

Diagnosis. A medium sized (40–44 mm SVL), shortlegged, shallow-headed, arboreal Cryptoblepharus, distinguished from Australian congeners by combination of modal values of five supraciliary scales, 24 mid-body scale rows and 50 paravertebral scales; mean head depth of 39.3% of head length; plain, rounded, dark pigmented plantar scales, and narrow, pale dorsolateral stripes.

Sternfeld diagnosed the taxon by “..b) Schuppen in 22–24 Reihen. Oberseite schhön rotbraun, die Dorso- lateralstreifen silberweiß, auf dem Halse zu einer dünnen Linie verschmälert, nach hinten zu allmählich wieder bre- iter werdend; die hellen Streifen mehr oder weniger scharf schwarz begrenzt. Schnauze außergewöhnlich kurz”. (“ Scales in 22–24 rows. Upper side beautifully bay, dorsolat- eral stripes silver, narrowing on the neck to a thin painted line, gradually becoming wide again on the back; the light stripes more or less keenly black edged. Snout exception- ally short”).

Diagnosis [clarus]: Very similar to conspecific C. p. pulcher but distinguished by having more midbody scale rows (mean 24.6 instead of 23.2), fourth finger (mean 15.1 instead of 14.6) and toe subdigital lamellae (mean 18.9 instead of 18.1), plantar scales (mean 9.8 instead of 8.9) and generally broader pale laterodorsal stripes. In addition, C. p. clarus has an allopatric distribution apparently restricted to far southern regions of Western and South Australia and may be further distinguished by two sexually dimorphic characters, where male C. p. clarus are larger than those of C. p. pulcher (mean SVL 36.1 instead of 33.7 mm) and female C. p. clarus have more paravertebral scales than those of C. p. pulcher (mean 50.2 instead of 48.4). With no fixed allelic differences, C. p. clarus is genetically similar to C. p. pulcher.

Diagnosis [pulcher]. Very similar to conspecific C. p. clarus but distinguished by having fewer midbody scale rows (mean 23.2 instead of 24.6), fourth finger (mean 14.6 instead of 15.1) and toe subdigital lamellae (mean 18.1 instead of 18.9), plantar scales (mean 8.9 instead of 9.8) and generally narrower pale laterodorsal stripes. In addition, C. p. pulcher has an allopatric distribution apparently restricted to eastern regions of Queensland and New South Wales and may be further distinguished by two sexually dimorphic characters, where male C. p. pulcher are smaller than those of C. p. clarus (mean SVL 33.7 instead of 36.1 mm) and female C. p. pulcher have fewer paravertebral scales than those of C. p. clarus (mean 48.4 instead of 50.2).

Cryptoblepharus suburbia was placed in the synonymy of C. virgatus by Shea
and Sadlier (1999), but was transferred to the synonymy of C. pulcher pulcher by HORNER 2007.
 
References
  • Bowles, F.D. 2000. A short note on the herpetofauna of Brisbane and its suburbs. Herpetological Bulletin (73): 27-29 - get paper here
  • Cogger H.G., Cameron EE & Cogger HM 1983. Zoological Catalogue of Australia, Volume 1: AMPHIBIA AND REPTILIA. Australian Government Publishing Service, Canberra.
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp.
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Cook, R. 1973. The wall lizard, Cryptoblepharus boutonii virgatus. Herpetofauna (Sydney) 6: 15-16
  • Covacevich, J. & Ingram, G. J. 1978. An undescribed species of rock dwelling Cryptoblepharus (Lacertilia: Scincidae). Memoirs of the Queensland Museum 18: 151-154
  • Daly, G., Pennay, M. and Gosper, C. 2001. Surveys of reptiles and amphibians at Razorback Nature Reserve, Keverstone State Forest and the Abercrombie Caves region of New South Wales. Herpetofauna (Sydney) 31(2): 82–91
  • Garman, S. 1901. Some reptiles and batrachians from Australasia. Bull. Mus. Comp. Zool. Harvard 39: 1-14 - get paper here
  • Greenbaum, E. 2000. Herpetofaunal observations in disparate habitats in south Sustralia, New South Wales, and queensland, Australia. Herpetological Bulletin (72): 6-16 - get paper here
  • Greer, A.E. 1974. The generic relationships of the scincid lizard genus Leiolopisma and its relatives. Australian Journal of Zoology 31: 1-67. - get paper here
  • Kay, G.M.; D. Michael; M. Crane; S. Okada; C. MacGregor; D. Florance; D. Trengove; L. McBurney; D. Blair; D.B. Lindenmayer. 2013. A list of reptiles and amphibians from Box Gum Grassy Woodlands in south-eastern Australia. Check List 9 (3):476-481 - get paper here
  • Mertens,R. 1931. Ablepharus boutonii (DESJARDIN) und seine geographische Variation. Zool. Jahrb. Syst. 61: 63-210
  • Mo, Matthew 2015. Herpetofaunal community of the constructed Lime Kiln Bay Wetland, south Sydney, New South Wales. Victorian Naturalist 132 (3): 64-72 - get paper here
  • Sternfeld, R. 1918. Zur Tiergeographie Papuasiens und der pazifischen Inselwelt. Abh. senckenb. naturf. Ges. (Frankfurt) 36: 375-436 - get paper here
  • Storr, G. M. 1961. Ablepharus boutonii clarus, a new skink from the Esperance District, Western Australia. Western Australian Naturalist 7: 176-178
  • Storr, G. M. 1976. The genus Cryptoblepharus (Lacertilia: Scincidae) in Western Australia. Rec. West. Aust. Mus. 4: 53-63 - get paper here
  • Storr, G. M., Smith, L. A. & Johnstone, R. E. 1981. Lizards of Western Australia. I. Skinks. Perth: University of Western Australia Press and Western Australian Museum, 200 pp.
  • Storr, G. M.; L. A. Smith, and R. E. Johnstone 1999. Lizards of Western Australia. I. Skinks. Revised Edition. Western Australian Museum
  • Wells, R. W. and Wellington, C. R. 1985. A classification of the Amphibia and Reptilia of Australia. Australian Journal of Herpetology, Supplementary Series, (1):1-61. - get paper here
  • Wells, R. W., and C. R. Wellington. 1989. A checklist of the amphibians and reptiles known from the Cumberland Plain Region, Sydney Basin, New South Wales, Australia. Australian Herpetologist, (506) :1-34
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
  • Wilson, Stephen K. & Knowles, David G. 1988. Australia's Reptiles: A Photographic Reference to the Terrestrial Reptiles of Australia. Cornstalk Publishing, Pymble, NSW, 447 pp.
  • Worrell, E. 1963. Reptiles of Australia. Angus & Robertson (Sydney), xv + 207 pp
 
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