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Ctenophorus maculatus (GRAY, 1831)

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Higher TaxaAgamidae (Amphibolurinae), Sauria, Iguania, Squamata (lizards)
Subspecies 
Common NamesE: Spotted Sand-dragon, Spotted Dragon 
SynonymUromastyx maculatus GRAY 1831
Grammatophora Gaimardii DUMÉRIL & BIBRON 1837: 470
Amphibolurus maculatus — BOULENGER 1885: 381
Amphibolurus maculatus — KINGHORN 1924
Amphibolurus maculatus — LICHT et al. 1966
Amphibolurus maculatus — COGGER 1983
Phthanodon maculatus — WELLS & WELLINGTON 1985: 19
Ctenophorus maculatus — COGGER 2000: 317
Ctenophorus maculatus — WILSON & SWAN 2010
Ctenophorus maculatus — CHAPPLE et al. 2019: 63
Ctenophorus maculatus — EDWARDS & HUTCHINSON 2023 
DistributionAustralia (Western Australia: west coast south of Shark Bay)

Type locality: unknown, but probably Shark Bay, W. A. (fide COGGER 1983)  
Reproductionoviparous 
TypesHolotype: MNHN-RA 1259 (considered as nomen nudum by Brygoo agamid type catalog) 
DiagnosisDiagnosis: Sandy to dark rust red dorsally, depending on location; relatively narrow middorsal zone because of extensive black dorsal margins of the wide, pale dorsolateral stripe (yellow in males) that overlies narrowly separated large black dorsolateral blotches; male with a distinct black throat chevron and rectangular chest badge (Figs. 4D, 5A–B, S8I–J, S9A). Femoral pore row reaches the knee. (Edwards & Hutchinson 2023) 
CommentGray's description referred to Cuvier's ms name applied to a specimen, almost certainly MNHP 1259, on which the name Grammatophora gaimardii is also based (COGGER 1983).

Habitat: shrubs, hummock grass.

Subspecies: all subspecies have been elevated to full species status by EDWARDS & HUTCHINSON 2023.

Distribution: see map in EDWARDS & HUTCHINSON 2023: 177 (Fig. 1) 
EtymologyFrom the Latin maculatus (spotted). (G. Shea, pers. comm., 9 Feb 2024) 
References
  • Boulenger, G.A. 1885. Catalogue of the lizards in the British Museum (Nat. Hist.) I. Geckonidae, Eublepharidae, Uroplatidae, Pygopodidae, Agamidae. London: 450 pp. - get paper here
  • Brygoo, E.R. 1988. Les types d’Agamidés (Reptiles, Sauriens) du Muséum national d’Histoire naturelle Catalogue critique. Bull. Mus. Nat. Hist. Nat. 10 (ser. 4) A (3), suppl.: 1-56
  • Chapple, David G.; Reid Tingley, Nicola J. Mitchell, Stewart L. Macdonald, J. Scott Keogh, Glenn M. Shea, Philip Bowles, Neil A. Cox, John C. Z. Woinarski 2019. The Action Plan for Australian Lizards and Snakes 2017. CSIRO, 663 pp. DOI: 10.1071/9781486309474 - get paper here
  • Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here
  • Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp.
  • Doughty, P.; Maryan, B.; Melville, J. & Austin, J. 2007. A new species of Ctenophorus (Lacertlia: Agamidae) from Lake Disappointment, Western Australia. Herpetologica 63 (1): 72-86 - get paper here
  • Duméril, A. M. C. and G. Bibron. 1837. Erpétologie Générale ou Histoire Naturelle Complete des Reptiles. Vol. 4. Libr. Encyclopédique Roret, Paris, 570 pp. - get paper here
  • Edwards, D., & Hutchinson, M. 2023. Sand Dragons: Species of the Ctenophorus maculatus Complex (Squamata: Agamidae) of Australia's Southern and Western Interior. Journal of Herpetology 57 (2): 176-196 - get paper here
  • Ellis, Ryan J. 2019. An annotated type catalogue of the dragon lizards (Reptilia: Squamata: Agamidae) in the collection of the Western Australian Museum. Records of the Western Australian Museum 34: 115–132 - get paper here
  • Escoriza Boj, D. 2005. Australia. Reptiles and Amphibians, Part 2: Desert and tropical savanna. Reptilia (GB) (41): 52-57 - get paper here
  • Glauert, L. 1959. Herpetological Miscellanea. XI. Dragon Lizards of the Genus Amphibolurus. Western Australian Naturalist 7 (2): 42-51 - get paper here
  • Gray, J. E. 1831. A synopsis of the species of Class Reptilia. In: Griffith, E & E. Pidgeon: The animal kingdom arranged in conformity with its organisation by the Baron Cuvier with additional descriptions of all the species hither named, and of many before noticed [V Whittaker, Treacher and Co., London: 481 + 110 pp. [1830] - get paper here
  • Kinghorn, J. Roy 1924. Reptiles and batrachians from south and south-west Australia. Rec. Austral. Mus. 14 (3): 163-183 - get paper here
  • Licht, P., W.R. Dawson, V.H. Shoemaker & A.R. Main 1966. Observations on the thermal relations of Western Australian lizards Copeia 1966 (1): 97-110. - get paper here
  • Melville, Jane; Schulte, James A.; Larson, Allan 2001. A molecular phylogenetic study of ecological diversification in the Australian lizard genus Ctenophorus. Journal of Experimental Zoology: Molecular and Developmental Evolution 291(4): 339-353 - get paper here
  • Sternfeld, R. 1925. Beiträge zur Herpetologie Inner-Australiens. Abhandlungen Herausgegeben von der Senckenbergischen Naturforschenden Gesellschaft, 38: 221—251
  • Storr, G. M. 1965. The Amphibolurus maculatus species-group (Lacertilia: Agamidae) in Western Australia. J R. Soc. West. Aust. 48: 45-54
  • Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
 
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