Cyclemys enigmatica FRITZ, GUICKING, AUER, SOMMER, WINK & HUNDSDÖRFER, 2008

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Higher Taxa	Geoemydidae (Geoemydinae), Testudinoidea, Testudines (turtles)
Subspecies
Common Names
Synonym	Cyclemys enigmatica FRITZ, GUICKING, AUER, SOMMER, WINK & HUNDSDÖRFER 2008 Cyclemys enigmatica — CHAN-ARD et al. 2015 Cyclemys enigmatica — TTWG 2021
Distribution	Indonesia (Sumatra), Southern Malay Peninsula and Greater Sunda Islands Type locality: Padang, Sumatra
Reproduction	oviparous.
Types	Holotype: NMW 9811 (Naturhistorisches Museum Vienna = NHMW), coll. Consul J. Schild 1901, stuffed adult female).
Diagnosis	Diagnosis: Adult: Dark coloured, with mainly or entirely dark brown to black shell. Carapace ovoid. Femoral mid-seam longer than or equal to anal mid-seam; anal notch wide and obtuse- angled. Young adults with dense black radiating pattern on carapace and plastron; aged adults often with uniform dark brown carapace and black plastron. Head and neck stripes absent. Crown of the head uniform brown, in life distinctly lighter coloured than temporal region. Palate and tongue in alcohol pure white. Hatchling (Fig. 9): Underside of the shell brownish-yellow, mottled with small darker brown specks; head and neck uniform dark, without stripes. For maximum size and com- parison with diagnostic characters of other species, see Table 3 in FRITZ et al. 2008: 10.
Comment	Similar species: has been confused with Heosemys grandis (e.g. in CHAN-ARD et al. 2015; the species in their Fig 2. is not C. enigmatica but Heosemys grandis, U. Fritz, pers. comm., 17. June 2012; Hikida 2015). Habitat: freshwater (Rivers, swamps)
Etymology	The species name enigmatica, Latinized adjective from the Greek noun aínigma (enigma, conundrum or riddle) refers to the long-lasting confusion caused by the distinct morphology of the new species and its introgressed mitochondrial genome.
References	Baker, Nick 2016. Dark-bellied Leaf Terrapin Cyclemys enigmatica at Pulau Langkawi, Peninsular Malaysia. SEAVR 2016: 1 - get paper here Bour, R. 2008. Global diversity of turtles (Chelonii; Reptilia) in freshwater. Hydrobiologia 595:593–598 - get paper here Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here Chan-ard, Tanya; Siriporn Thong-aree, Michael Cota and Sunchai Makchai 2011. Cyclemys of the Peat Swamp of Southernmost Thailand. The Thailand Natural History Museum Journal 5(2): 107-112 Fritz, U.; D Guicking, M Auer, R. S Sommer, M Wink, A. K Hundsdörfer 2008. Diversity of the Southeast Asian leaf turtle genus Cyclemys: how many leaves on its tree of life? Zoologica Scripta doi:10.1111/j.1463-6409.2008.00332. - get paper here Gemel, R.; G. Gassner & S. Schweiger 2019. Katalog der Typen der Herpetologischen Sammlung des Naturhistorischen Museums Wien – 2018. Ann. Naturhist. Mus. Wien, B 121: 33–248 Hikida, Tsutomu 2015. Book Review: A Field Guide to the Reptiles of Thailand. Nat. Hist. Bull. Siam Soc. 61(1): 41–51 TTWG; Rhodin, A.G.J., Iverson, J.B., Bour, R., Fritz, U., Georges, A., Shaffer, H.B., and van Dijk, P.P. 2021. Turtles of the World: Annotated Checklist and Atlas of Taxonomy, Synonymy, Distribution, and Conservation Status (9th Ed.). In: Rhodin, A.G.J., Iverson, J.B., van Dijk, P.P., Stanford, C.B., Goode, E.V., Buhlmann, K.A., and Mittermeier, R.A. (Eds.). Chelonian Research Monographs 8:1–472. doi:10.3854/crm.8.checklist.atlas.v9.2021. - get paper here
External links	IUCN Red List Google images

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