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Cyrtodactylus bokorensis MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymCyrtodactylus bokorensis MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER 2019
Gymnodactylus intermedius SMITH 1935: 44
Cyrtodactylus intermedius STUART & EMMETT 2006: 17 
DistributionCambodia (Kampot)

Type locality: Bokor National Park Field Station, Kampot Province, Cambodia (10 37.611'N, 104 01.541'E); 1033 m elevation  
Reproduction 
TypesHolotype: LSUHC 8554, adult male, collected on 10 July 2007 by L. Lee Grismer and Neang Thy. Paratypes. Adult males LSUHC 8541–42, adult females LSUHC 8548–50 and 8555 bear the same collection data as the holotype. 
DiagnosisDiagnosis. Adult males reaching 81.9 mm SVL, adult females reaching 93 mm SVL; eight supralabials, 8–10 infralabials; 30–33 paravertebral tubercles; 18–20 longitudinal rows of dorsal tubercles; 42–48 rows of ventral scales; six or seven expanded subdigital lamellae proximal to the digital inflection, 11–14 unmodified, distal, subdigital lamellae; 18–20 total subdigital lamellae on fourth toe; enlarged femoral and precloacal scales vary from being separated by a diastema to continuous; 26–30 enlarged femoral scales; proximal femoral scales less than half the size of distal proximal scales; 7–10 enlarged precloacal scales with pores on each in males; two or three rows of enlarged post-precloacal scales; two or three postcloacal tubercles; no pocketing present between digits of hind or forefeet; dark pigmented blotches on top of head vary from present to absent; posterior border of nuchal loop pointed, chevron-shaped; and four or five dark body bands (summarized in Tables 7 + 10 in Murdoch et al. 2019).

Comparisons. Cyrtodactylus bokorensis sp. nov. is a member of the eastern group and the sister species of Cyrtodactylus laangensis sp. nov. from Phnom Laang from which it is separated by 3.9% sequence divergence (Table 4 in Murdoch et al. 2019). The PCA analysis indicates Cyrtodactylus bokorensis sp. nov. is distinct in morphospace from all other species with the exceptions of C. septimontium sp. nov. and C. cardamomensis sp. nov. with which there is overlap (Fig 6). The concatenated DAPC analysis Cyrtodactylus bokorensis sp. nov. shows complete separation from all other species (Fig. 7). Cyrtodactylus bokorensis sp. nov. is well-differentiated from all other members of the C. intermedius complex by having varying combinations of statistically different mean values of supralabial and infralabial scales, paravertebral tubercles, longitudinal rows of tubercles, ventral scales, unmodified, expanded, and total number of subdigital lamellae, enlarged femoral scales, precloacal scales, and postcloacal tubercles (Table 6). It differs further from all other members of the complex in the posterior border of the nuchal loop being chevron shaped and relatively pointed as opposed to rounded. It is differentiated from C. thylacodactylus sp. nov. in lacking interdigital pocketing (Table 7). 
CommentHabitat: scattered sandstone rocks and boulders among highly disturbed wind-blown scrub vegetation. Also abundant at 350 m elevation in wet evergreen forest associated with rocky areas on the southern side of Bokor NP and therefore probably common throughout Bokor National Park. 
EtymologyThe specific epithet, bokorensis sp. nov., is an adjective in reference to type locality of the Bokor Plateau to which it is presumably endemic. 
References
  • MURDOCH, MATTHEW L.; L. LEE GRISMER, PERRY L. JR. WOOD, THY NEANG, NIKOLAY A. POYARKOV, NGO VAN TRI, ROMAN A. NAZAROV, ANCHALEE AOWPHOL, OLIVIER S.G. PAUWELS, HUNG NGOC NGUYEN, JESSE L. GRISMER 2019. Six new species of the Cyrtodactylus intermedius complex (Squamata: Gekkonidae) from the Cardamom Mountains and associated highlands of Southeast Asia. Zootaxa 4554 (1): 001–062 - get paper here
  • Smith,M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Stuart, B.L. & Emmett, D.A. 2006. A Collection of Amphibians and Reptiles from the Cardamom Mountains, Southwestern Cambodia. Fieldiana Zool. N.S. (109): 1-27 - get paper here
 
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