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Cyrtodactylus dammathetensis GRISMER, WOOD JR, THURA, ZIN, QUAH, MURDOCH, GRISMER, LIN, KYAW & LWIN, 2017

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesDammathet cave bent-toed gecko 
SynonymCyrtodactylus dammathetensis GRISMER, WOOD JR, THURA, ZIN, QUAH, MURDOCH, GRISMER, LIN, KYAW & LWIN 2017: 62 
DistributionMyanmar (Mawlamyine District, Mon State)

Type locality: Dammathet Cave 19.8 km east of Mawlamyine, Mawlamyine District, Mon State, Myanmar (N16°30.380, E97°48.629; 25 m in elevation).  
Reproduction 
TypesHolotype: LSUHC 12862, Adult male collected on 7 October 2016 at 2000 h by Matthew L. Murdoch, L. Lee Grismer, Marta S. Grismer, Myint Kyaw Thura, Evan S. H. Quah, Perry L. Wood, Jr., Thaw Zin and Htet Kyaw.
Paratypes: Adult females LSUHC 12863–64 bear the same collection data as the holotype. 
DiagnosisDiagnosis: Cyrtodactylus dammathetensis sp. nov. differs from all congeners by having the unique combination of nine supralabials; seven or eight infralabials; 13–15 longitudinal rows of body tubercles; 31–33 paravertebral tubercles; 25–28 ventral scales; relatively long digits with eight or nine expanded subdigital lamellae proximal to the digital inflection on the fourth toe, 12 or 13 unmodified distal subdigital lamellae,21 total subdigital lamellae;raised,moderately keeled, dorsal body tubercles; tubercles extend beyond base of tail; enlarged femoral and precloacal scales continuous; enlarged proximal femoral scales less than one-half the size of enlarged distal femoral scales; 31–36 enlarged femoral scales; 36 femoral pores in males; eight or nine enlarged precloacal scales; nine precloacal pores in males; three rows of enlarged post- precloacal scales; median subcaudal scales three times as wide as long, extending to lateral surface of tail; top of head darkly mottled, no yellow reticulum; nuchal loop variably paired, no pronounced, anterior, azygous notch, posterior border jagged; five jagged, dark, dorsal bands with paravertebral elements, wider than interspaces, lacking lightened centres, not edged with white tubercles; nape band present; dark markings in dorsal interspaces; ventrolateral folds faintly whitish; anterodorsal margins of thighs and brachia darkly pigmented; ten or 11 light caudal bands variably bearing dark markings, not encircling tail; and ten or 11 dark caudal bands wider than light caudal bands.

Comparisons: Cyrtodactylus dammathetensis sp. nov. is part of the sinyineensis group. The PCA and DAPC analyses indicate that the species of this group are completely separate in morphospace where the first two principal components account for 63% of the total variation (Fig. 12) and load most heavily for numbers of infralabials, longitudinal rows of tubercles, expanded subdigital lamellae on the fourth toe and post-precloacal scale rows (Table S3). Cyrtodactylus dammathetensis sp. nov. is well-differentiated from C. aequalis, C. sinyineensis sp. nov. and C. welpyanensis sp. nov. by having varying combinations of statistically different mean values of infralabial scales, ventral scales, enlarged femoral scales, post-preprecloacal scale rows, longitudinal rows of body tubercles, precloacal scales and body bands (Table 3). It differs further from C. sinyineensis sp. nov. and C. welpyanensis sp. nov. in having paravertebral elements in the dorsal banding pattern (Table 8). Morphological and colour pattern differences from other species in the Indo-Chinese clade are listed in Table 8. Genetic distances among the species of this group range from 11.0 to 16.5% (Table 10). 
Comment 
EtymologyThe specific epithet, dammathetensis, is a noun in apposition in reference to the type locality of Dammathet Cave. 
References
  • Grismer, L. Lee; Perry L. Wood Jr., Myint Kyaw Thura, Evan S.H. Quah, Matthew L. Murdoch, Marta S. Grismer, Mark W. Herr, Aung Lin & Htet Kyaw 2018. Three more new species of Cyrtodactylus (Squamata: Gekkonidae) from the Salween Basin of eastern Myanmar underscore the urgent need for the conservation of karst habitats. Journal of Natural History, 52:19-20, 1243-1294 - get paper here
  • Grismer, L.L.; PERRY L. WOOD, JR., MYINT KYAW THURA, THAW ZIN, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MARTA S. GRISMER, AUNG LIN, HTET KYAW and NGWE LWIN 2017. Twelve new species of Cyrtodactylus Gray (Squamata: Gekkonidae) from isolated limestone habitats in east- central and southern Myanmar demonstrate high localized diversity and unprecedented microendemism Zoological Journal of the Linnean Society 182: 862-959
 
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