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Cyrtodactylus himalayicus (ANNANDALE, 1906)

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesHimalaya bent-toed Gecko 
SynonymGymnodactylus himalayicus ANNANDALE 1906
Cyrtodactylus himalayicus — AGARWAL et al. 2018 
DistributionIndia (West Bengal, Darjeeling

Type locality: “Kurseong, Darjeeling dist. (5,000 ft)” [=Kurseong subdivision (ca. 1524 m elevation), Darjeeling district, West Bengal state, India]  
Reproductionoviparous 
TypesHolotype: ZSI (ZSIK) 15716, male, collected by N. Annandale, ca. 1906 (Annandale 1906). 
DiagnosisDiagnosis and comparisons with regional congeners. A small sized Cyrtodactylus, snout-vent length to 64.5 mm; body relatively slender; limbs and digits moderately long; two or three pairs of well-developed postmentals, inner pair longer than outer two and in broad contact behind mental; 19–21 rows of smooth weakly raised to bluntly conical, rounded dorsal tubercles; 33 or 34 [35 count in Annandale 1906] ventral scales between outermost ventral scale rows enlarged relative to granular flank scales; ventrolateral folds absent; no precloacal groove; 10 precloacal pores in a single series on the male and an equivalent 10 fine pores/ pitted scales on the female, femoral pores absent on both sexes; scales bordering precloacal pore row are approximately 3x enlarged relative to pore- bearing scales. Nine basal subdigital lamellae and 11 distal lamellae on Digit IV of pes [n=1: toe tips broken off on ZSIK 15716]. Subcaudal scalation of original tail without single median series of enlarged plates, though subcaudals are moderately enlarged relative to dorsal caudal scales. Dorsal pattern of approximately eight paired transversely arranged small dark spots on trunk and sacrum of male, or female with four almost continuous longitudinal brown stripes on trunk. Original tail with approximately 13 dark bands.

Cyrtodactylus himalayicus comb. nov. differs from the following species in the presence of only precloacal pores on males (vs. PcP + femoral pores): C. annandalei Bauer, C. fasciolatum, C. gubernatoris, C. russelli and C. slowinskii. Cyrtodactylus himalayicus comb. nov. differs from the following species by possessing 10 precloacal pores: C. brevidactylus Bauer (8 PcP), C. cayuensis (6–9 PcP), C. mandalayensis (8 PcP + single enlarged pore- bearing scale posterior to the series), C. markuscombaii (7 PcP), C. martinstolli (0–8 PcP); and from the following species by possessing precloacal pores only (vs. precloacofemoral pores): C. gansi (16–29 PcFP), and C. tamaiensis (40 PcFP). Cyrtodactylus himalayicus comb. nov. differs from the following species by having 19–21 dorsal tubercle rows: C. brevidactylus (27 DTR), C. mandalayensis (18 DTR), C. markuscombaii (14–15 DTR), and C. wakeorum Bauer (24 DTR). Cyrtodactylus himalayicus comb. nov. lacks a ventrolateral fold, which distinguishes it from C. ayeyarwadyensis, C. cayuensis, C. mandalayensis and C. martinstolli. Cyrtodactylus himalayicus comb. nov. differs from C. khasiensis by having enlarged plate-like scales posteriorly bordering the pore-bearing scale series (equivalent scales not enlarged on C. khasiensis), and snout to vent length to 64.5 mm (to 83.3 mm for C. khasiensis). Refer to the diagnosis section of new taxa for comparisons with Cyrtodactylus himalayicus comb. nov. 
Comment 
Etymology 
References
  • AGARWAL, ISHAN; STEPHEN MAHONY, VARAD B. GIRI, R. CHAITANYA, AARON M. BAUER 2018. Two new species of bent toed geckos, Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Northeast India with comments on name-bearing types from the region. Zootaxa 4420 (3): 334-356 - get paper here
  • AGARWAL, ISHAN; STEPHEN MAHONY, VARAD B. GIRI, R. CHAITANYA, AARON M. BAUER 2018. Six new Cyrtodactylus (Squamata: Gekkonidae) from northeast India Zootaxa 4524 (5): 501–535 - get paper here
  • Annandale, N. 1906. A new gecko from the eastern Himalayas. J. Proc. Asiat. Soc. Bengal 2: 287—288 - get paper here
 
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