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Cyrtodactylus khammouanensis NAZAROV, POYARKOV, ORLOV, NGUYEN, MILTO, MARTYNOV, KONSTANTINOV & CHULISOV, 2014

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies 
Common NamesE: Khammouane Bent-toed Gecko 
SynonymCyrtodactylus khammouanensis NAZAROV, POYARKOV, ORLOV, NGUYEN, MILTO, MARTYNOV, KONSTANTINOV & CHULISOV 2014 
DistributionLaos (Khammouane)

Type locality: environs of Na Phao Village, Boulapha Dis- trict, Khammouane Province, Laos (17°34 ́57.1 ́ ́N, 105°44 ́37.3 ́ ́E, elevation 170 m a.s.l.).  
Reproductionoviparous (manual imputation, fide Zimin et al. 2022) 
TypesHolotype: ZMMU R-13982, Adult male, (field number FN 192) collected on 16 June 2009 by Nikolai Orlov, Sang Ngoc Nguyen and Konstantin Milto. Paratypes. One adult male ZIN 28250 (FN 193) and two adult females ZMMU R-13983 (FN 257); ZIN 28251 (FN 191). All type specimens are with the same data as the holotype. 
DiagnosisDiagnosis: Relatively small Cyrtodactylus with a maximum SVL of 73 mm; the new species is distinguished from all other congeners by the combination of the following morphological characters. Dorsal pattern consisting of wide dark nuchal band and 4–5 dark wide transverse bands with smooth edges. Widths of interspaces between dorsal bands are less than widths of the bands. Dorsal surface of head is light yellowish, without any clear pattern. Dark wide transverse bands present on the tail. Roundish weakly keeled tubercles present on the dorsum, hind limbs, temporal region of the head, on the tail base and absent on the dorsal surface of the head. Ventrals in 32–38 longitudinal rows at midbody. Lateral folds weakly developed. Continuous series of 40–44 pre- cloacal and femoral pores in males reach the bend of knee. Enlarged femoral scales present, 5–6 enlarged postcloacal spurs in both sexes. Subcaudal scales transversally enlarged in one median row. 
CommentAbundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet of the new species is derived from the name of Khammouane Province of Laos, renowned for its great diversity of Cyrtodactylus geckoes. 
References
  • LUU, VINH QUANG; THOMAS CALAME, Truong QUang Nguyen, MICHAEL BONKOWSKI & THOMAS ZIEGLER 2015. A new species of Cyrtodactylus (Squamata: Gekkonidae) from the limestone forest of Khammouane Province, central Laos. Zootaxa 4058 (3): 388–402 - get paper here
  • LUU, VINH QUANG; Truong QUang Nguyen, MINH DUC LE, MICHAEL BONKOWSKI, THOMAS ZIEGLER 2016. A new species of karst-dwelling bent-toed gecko (Squamata: Gekkonidae) from Khammouane Province, central Laos. Zootaxa 4079 (1): 087–102 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Nazarov, R.A.; N.A. Poyarkov Jr., N.L. Orlov, N.S. Nguyen, K.D. Milto, A.A. Martynov, E.L. Konstantinov and A.S. Chulisov 2015. A review of genus Cyrtodactylus (Reptilia: Sauria: Gekkonidae) in fauna of Laos with description of four new species. Труды Зоологического института РАН 318 (4): 391–423 [Proceedings of the Zoological Institute RAS 318(4):391-423, 2014]
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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