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Cyrtodactylus lenggongensis GRISMER, WOOD, ANUAR, GRISMER, QUAH, MURDOCH, MUIN, DAVIS, AUILAR, KLABACKA, COBOS, AOWPHOL & SITES, 2016

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesEnglish: Lenggong Bent-toed Gecko
Malay: Cicak Jari-bengkok Lenggong 
SynonymCyrtodactylus lenggongensis GRISMER, WOOD, ANUAR, GRISMER, QUAH, MURDOCH, MUIN, DAVIS, AUILAR, KLABACKA, COBOS, AOWPHOL & SITES 2016
Cyrtodactylus bintangrendah — GRISMER et al. 2012: 32
Cyrtodactylus bintangrendah — GRISMER et al. 2014: 360 
DistributionPeninsular Malaysia (Perak)

Type locality: Lenggong Valley, Perak, Peninsular Malaysia (5°06.431” N 100°58.322” E; at 104 m elevation.  
Reproductionoviparous. Of the four adult females sampled (LSUHC 12159, 12162, 12173–74), none were gravid although one hatchling was collected (LSUHC 12177). This would suggest that C. hidupselamanya sp. nov. does not breed year-round and that April is at the end of its reproductive season (Grismer et al. 2016). 
TypesHolotype. LSUHC 9974, Adult male, collected on 29 October 2011 by E.S.H. Quah, Fatim, S.B.M., and Nor Amira, B.A.R.
Paratypes. Paratypes, LSUHC 9975–77 bear the same collection data as the holotype. These were all once type material of Cyrtodactylus bintangrendah but are now excluded from that species. 
CommentDiagnosis. Cyrtodactylus lenggongensis sp. nov. can be differentiated from all other species of Cyrtodactylus by having a maximum SVL of 103.1 mm; 10 or 11 supralabials; 8–10 infralabials; prominent tuberculation on body; no tubercles on ventral surface of forelimbs, gular region, or in ventrolateral body folds; tubercles on anterior one-third of tail; 36–41 paravertebral tubercles; 22–25 longitudinal tubercle rows; 32 or 33 ventral scales; 20–23 subdigital lamellae on fourth toe; no femoral pores; 39–41 femoroprecloacal scales; deep precloacal groove; four or five dark dorsal body bands; body band width less than to slightly more than interspace width; body bands and nuchal loop edged with a thin yellowish, tubercle-bearing lines; no scattered white tubercles on dorsum; 14 dark caudal bands on original tail; and light-colored caudal bands immaculate. These characters are scored across all species of the C. pulchellus complex in Table 5 (in Grismer et al. 2016).

Comparisons. Cyrtodactylus lenggongensis sp. nov. is differentiated from all other species of the C. pulchellus complex by having a combination of low, rounded tubercles on body; no tubercles on ventral surfaces of forelimbs, gular region or in the ventrolateral body folds; 36–41 paravertebral tubercles; 32 or 33 ventral scales; 39–41 femoroprecloacal pores; a deep precloacal groove; four or five body bands that are less than to slightly wider than the width of the interspace; body bands and nuchal loop edged with yellowish tubercles; and no scattered white tubercles on dorsum (Table 5). Within the C. pulchellus complex, C. lenggongensis sp. nov. is the sister species of C. bintangrendah sensu stricto to which it was once considered conspecific (Grismer et al. 2012, 2014a). However it differs from C. bintangrendah sensu stricto in having weaker body tuberculation; no tubercles in the ventrolateral folds; tubercles on forelimbs and legs are low, rounded and widely separated versus being large and in near contact; fewer femoroprecloacal scales (39–41 versus 41–46); 14 versus eight or nine dark caudal bands on the original tail, caudal bands encircling tail forming rings; and a smaller maximum SVL (103.1 versus 114.4).

Habitat: Cyrtodactylus lenggongensis sp. nov. is saxicolous and found on limestone karst outcrops and in the vicinity of the mouths of limestone caves (Fig. 7). During the day, geckos are occasionally observed hiding in cracks and crevices of the outcrops. At night, lizards emerge and can be found perching or crawling over the walls of the karst formation 1–3 m above the ground. No lizards were found on the surrounding vegetation (Grismer et al. 2016). 
EtymologyThe specific epithet lenggongensis refers to the Lenggong Valley, Perak where this species is endemic. 
References
  • GRISMER, L. LEE; PERRY L. WOOD, JR, SHAHRUL ANUAR, MARTA S. GRISMER, EVAN S. H. QUAH, MATTHEW L. MURDOCH, MOHD ABDUL MUIN, HAYDEN R. DAVIS, CÉSAR AGUILAR, RANDY KLABACKA, ANTHONY J. COBOS, ANCHALEE AOWPHOL, JACK W. SITES, JR 2016. Two new Bent-toed Geckos of the Cyrtodactylus pulchellus complex from Peninsular Malaysia and multiple instances of convergent adaptation to limestone forest ecosystems. Zootaxa 4105 (5): 401-429 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, JR., EVAN S. H. QUAH, SHAHRUL ANUAR, MOHD. ABDUL MUIN, MONTRI SUMONTHA, NORHAYATI AHMAD, AARON M. BAUER, SANSAREEYA WANGKULANGKUL, JESSE L. GRISMER & OLIVIER S. G. PAUWELS 2012. A phylogeny and taxonomy of the Thai-Malay Peninsula Bent-toed Geckos of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae): combined morphological and molecular analyses with descriptions of seven new species. Zootaxa 3520: 1–55 - get paper here
  • GRISMER, L. LEE; PERRY L. WOOD, SHAHRUL ANUAR, EVAN S. H. QUAH, MOHD ABDUL MUIN, MAKETAB MOHAMED, CHAN KIN ONN, ALEXANDRA X. SUMARLI, ARIEL I. LOREDO, HEATHER M. HEINZ 2014. The phylogenetic relationships of three new species of the Cyrtodactylus pulchellus complex (Squamata: Gekkonidae) from poorly explored regions in northeastern Peninsular Malaysia. Zootaxa 3786 (3): 359–381 - get paper here
 
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