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Cyrtodactylus limajalur DAVIS, BAUER, JACKMAN, NASHRIQ & DAS, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Five-banded Bent-toed Gecko 
SynonymCyrtodactylus limajalur DAVIS, BAUER, JACKMAN, NASHRIQ & DAS 2019 
DistributionEast Malaysia (Borneo: Sarawak)

Type locality: near Kampung Tubih Mawang, Serian, Sarawak, East Malaysia. (01°17'47.62"N; 110°26'33.00"E; 88 m elevation; WGS 1984)  
Reproduction 
TypesHolotype: CAS 262848, adult male (Fig. 5 in Davis et al. 2019), collected by Izneil Nashriq and Hayden Davis on August 06, 2017 at 20:00–22:00 hrs (Fig. 1).
Paratypes. Paratypes (CAS 262846; CAS 262847; and CAS262849) (Fig. 5) have the same collection data as the holotype. 
DiagnosisDiagnosis. Cyrtodactylus limajalur sp. nov. can be differentiated from all other species of Cyrtodactylus by the combination of the following characters: maximum SVL of at least 94 mm; 10–12 supralabials; 9–11 infralabials; weak tuberculation on body; no tubercles on ventral surface of forelimbs, gular region, or ventrolateral folds; 34–38 paravertebral tubercles; 11–13 longitudinal tubercle rows; 33–42 ventral scales; 19–22 subdigital lamellae on fourth toe; 5–6 enlarged femoral scales; no femoral pores; 7–8 precloacal pores; precloacal pit in males; enlarged median row of transverse scales; 5 dark dorsal body bands; body bands more than two times the width of interspaces; rostral chevron just posterior to orbitals; no white line edging the body bands and nuchal loop; and no scattered white tubercles on dorsum. These characters are scored across all currently recorded Bornean Cyrtodactylus species in Table 4 in Davis et al. 2019: 344.

Comparison. Cyrtodactylus limajalur sp. nov. differs from all its Bornean congeners by one or more morphological characteristics. The new species is distinguished from C. baluensis by having a distinct banding pattern as opposed to bands and blotches; it is distinguished from C. cavernicolus by having a lower number of ventral scales (33–42 versus 51–58), enlarged subcaudals, and a precloacal pit as opposed to a precloacal groove; it is distinguished from C. consobrinus by having a smaller adult maximum SVL (94 mm versus 125 mm), and no white reticulated pattern on occiput; it is distinguished from C. ingeri by having a precloacal pit as opposed to no depression, 5–6 enlarged femoral scales as opposed to none, and a distinct banding pattern as opposed to bands and blotches; it is distinguished from C. malayanus by having no tubercles on the upper arm, and a lower number of ventral scales (33–42 versus 58–62); it is distinguished from C. matsuii by having enlarged subcaudals, 5–6 enlarged femoral scales as opposed to none, and a smaller maximum SVL (94 mm versus 105 mm); it is distinguished from C. pubisulcus by having enlarged subcaudals, 5–6 enlarged femoral scales as opposed to none, and a precloacal pit as opposed to a precloacal groove; and it is distinguished from C. yoshii by having no tubercles on the upper arm, a lower number of ventral scales (33–42 versus 50–58), a lower number of subdigital lamellae (19–22 versus 25–30), and 5–6 enlarged femoral scales as opposed to none. 
CommentSympatry: Cyrtodactylus consobrinus and C. pubisulcus. However, C. consobrinus was seen almost exclusively on large tree trunks and C. pubisulcus was only found on low-lying vegetation, indicating that they may not be syntopic with C. limajalur sp. nov. 
EtymologyThe specific epithet limajalur is in reference to the banding pattern of the species. “Lima jalur” loosely translated from Malay is five-banded. Cyrtodactylus limajalur sp. nov. has a very distinct banding pattern that is consistently comprised of five bands. This species is the first Bornean Cyrtodactylus species that displays a consistent banding pattern, without blotches or other markings. 
References
  • DAVIS, HAYDEN R.; AARON M. BAUER, TODD R. JACKMAN, IZNEIL NASHRIQ & INDRANEIL DAS 2019. Uncovering karst endemism within Borneo: two new Cyrtodactylus species from Sarawak, Malaysia. Zootaxa 4614 (2): 331–352 - get paper here
 
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