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Cyrtodactylus septimontium MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER, 2019

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Bảy Núi (Seven Mountains) Bent-toed Gecko 
SynonymCyrtodactylus septimontium MURDOCH, GRISMER, WOOD, NEANG, POYARKOV, TRI, NAZAROV, AOWPHOL, PAUWELS, NGUYEN & GRISMER 2019: 46
Cyrtodactylus intermedius — NGO & GRISMER 2006: 328
Cyrtodactylus intermedius — BOBROV & SEMENOV 2008: 236
Cyrtodactylus intermedius — NGUYEN et al. 2009: 768
Cyrtodactylus intermedius — NGUYEN et al. 2013
Cyrtodactylus intermedius — NGUYEN et al. 2014 
DistributionVietnam (An Giang)

Type locality: Nui Cam Mountain, An Giang province, Vietnam  
Reproduction 
TypesHolotype: LSUHC 12484, adult male, collected on 5 December 2005 by Ngo Van Tri. Paratypes. Adult male LSUHC 8244 and adult female 8243 bear the same collection data as holotype. Adult female LSUHC 12485 was collected at the same location. Adult male LSUHC 12488 and adult females LSUHC 12486–87 were collected on the adjacent hill Ta Soc Mountain, An Giang province, Vietnam. Adult males NAP– 05274, 05277, and adult females NAP–05272–73, 05275–76, all registered as ZMMU R–14773, were collected at Nui Cam mountain, An Giang Province, Vietnam, on 26 August 2015 by N.A. Poyarkov and Nguyen Ngoc Hung (N 10.51169; E 104.97193; 436 meters in elevation). Adult males NAP–05320–21, 05323, 05325, and adult females NAP–05322, 05324, all registered as ZMMU R–14775, were collected at Nui Co To mountain, An Giang Province, Vietnam, on 25 August 2015 by N.A. Poyarkov and Nguyen Ngoc Hung (N 10.38607; E 104.99169; 388 meters in elevation). 
DiagnosisDiagnosis. Adult males reaching 85.5 mm SVL, adult females reaching 90.4 mm SVL; 8–10 supralabials, 9– 11 infralabials; 23–30 paravertebral tubercles; 16–20 longitudinal rows of dorsal tubercles; 37–46 rows of ventral scales; six or seven expanded subdigital lamellae proximal to the digital inflection, 11–13 unmodified, distal, subdigital lamellae; 17–20 total subdigital lamellae on fourth toe; enlarged femoral and precloacal scales continuous; 24–33 enlarged femoral scales; proximal femoral scales less than one-half the size of distal proximal scales; 6–8 enlarged precloacal scales with pores on each in males; two or three rows of enlarged post-precloacal scales; two postcloacal tubercles; interdigital pocketing absent; dark pigmented blotches absent on top of head; posterior border of nuchal loop rounded; four dark body bands (summarized in Tables 7 + 14 in Murdoch et al. 2019).

Comparisons. Cyrtodactylus septimontium sp. nov. is a member of the eastern group sister species to C. intermedius complex incertae sedis 2 from Kirirom National Park from which it is separated by 2.9% sequence divergence (Table 4 in Murdoch et al. 2019). The PCA analysis indicates C. septimontium sp. nov. is well-separated from all other species in morphospace except for overlap with C. bokorensis sp. nov. and C. intermedius (Figs. 6). DAPC analysis shows C. septimontium as distinct from C. bokorensis sp. nov., C. laangensis sp. nov., and C. thylacodactylus sp. nov. but having general overlap with the remaining members of the complex. C. septimontium sp. nov. is well-differentiated from all other members of the C. intermedius complex by having varying combinations of statistically different mean values of supralabial and infralabial scales, paravertebral tubercles, ventral scales, unmodified, expanded, and total number of subdigital lamellae, enlarged femoral scales, pre-cloacal scales, and post-cloacal tubercles (Table 6). It differs from C. auralensis sp. nov. by lacking dark blotches on the top of the head. It differs from C. bokorensis sp. nov. in having a nuchal loop with a rounded posterior border. It differs from C. laangensis in having contact between femoral and precloacal scales. It differs from C. cardamomensis and C. thylacodactylus sp. nov. in having proximal femoral scales that are less than half the size of the distal scales. It is further differentiated from C. thylacodactylus sp. nov. in lacking interdigital pocketing (Table 7). 
CommentHabitat: large granite rocks, occasionally on tree trunks up to 2.5 m above the ground, or on the forest floor hiding in the leaf litter. Most specimens were recorded on vertical surfaces of large granite rocks or in caverns between them.

Sympatry: Cnemaspis nuicamensis, Gehyra mutilata, Hemiphyllodactylus sp., Calotes versicolor, Lipinia vittigera and Scincella cf. rufocaudata. On Nui Co To Mountain Cyrtodactylus septimontium sp. nov. was also recorded mostly on large granite rocks at elevations 350 – 450 m a.s.l., where it was found in sympatry with Cyrtodactylus grismeri and Cnemaspis tucdupensis. 
EtymologyThe specific epithet, septimontium, is a noun in apposition in reference to a Latinization of the Vietnamese name for the hill range Bảy Núi meaning the “Seven Mountains”. 
References
  • Bobrov V.V., Semenov D.V. 2008. Lizards of Vietnam [in Russian]. Moscow, 236 pp.
  • MURDOCH, MATTHEW L.; L. LEE GRISMER, PERRY L. JR. WOOD, THY NEANG, NIKOLAY A. POYARKOV, NGO VAN TRI, ROMAN A. NAZAROV, ANCHALEE AOWPHOL, OLIVIER S.G. PAUWELS, HUNG NGOC NGUYEN, JESSE L. GRISMER 2019. Six new species of the Cyrtodactylus intermedius complex (Squamata: Gekkonidae) from the Cardamom Mountains and associated highlands of Southeast Asia. Zootaxa 4554 (1): 001–062 - get paper here
  • Ngo, V.T. & Grismer, L.L. 2006. The first record of Cardamom slender–toed gecko – Cyrtodactylus intermedius Smith, 1917 in Vietnam. Proceedings of the 5th Scientific Conference of University of Natural Science, Ho Chi Minh City, 30 November 2006, 328
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • NGUYEN, SANG NGOC; JUN-XIAO YANG, THANH-NGAN THI LE, LUAN THANH NGUYEN, NIKOLAI L. ORLOV, CHUNG VAN HOANG, TRUONG QUANG NGUYEN, JIE-QIONG JIN, DING-QI RAO, THAO NGOC HOANG, JING CHE, ROBERT W. MURPHY, YA-PING ZHANG 2014. DNA barcoding of Vietnamese bent-toed geckos (Squamata: Gekkonidae: Cyrtodactylus) and the description of a new species. Zootaxa 3784 (1): 048–066 - get paper here
  • NGUYEN, SANG NGOC; THANH-NGAN THI LE, THI ANH DAO TRAN, NIKOLAI L. ORLOV, AMY LATHROP, ROSS D. MACCULLOCH, THUY-DUONG THI LE, JIE-QIONG JIN, LUAN THANH NGUYEN, TAO THIEN NGUYEN, DAT DUC HOANG, JING CHE, ROBERT W. MURPHY & YA-PING ZHANG 2013. Phylogeny of the Cyrtodactylus irregularis species complex (Squamata: Gekkonidae) from Vietnam with the description of two new species. Zootaxa 3737 (4): 399–414 - get paper here
 
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