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Cyrtodactylus tanahjampea RIYANTO, HAMIDY & MCGUIRE, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesE: Tanahjampea bent-toed gecko
Indonesian: Cicak Jari-lengkung Tanahjampea 
SynonymCyrtodactylus tanahjampea RIYANTO, HAMIDY & MCGUIRE 2018 
DistributionIndonesia (Sulawesi: Tanahjampea Island)

Type locality: Telkom tower road on Pasimasunggu district, Kepulauan Selayar regency, South Sulawesi Province, Tanahjampea Island, Indonesia (07°5’09.4” S, 120°39’29.2” E, 316 m above sea level  
Reproduction 
TypesHolotype: MZB Lace.5675 (Fig.2), adult male, collected by J. A. McGuire, Christopher J. Hayden, and Ted Townsend on 18 October 2005, 9:07 pm.
Paratype. MZB.Lace.5671–72, MZB.Lace.5674, adult females, vicinity of Makminasa, Pasimasunggu district, Kepulauan Selayar regency, South Sulawesi Province, Tanahjampea Island, Indonesia (07o05’04.9” S, 120o40’58.3” E, 195 m above sea level), collected by Christopher J. Hayden on 16 October 2005 11:00 pm; MZB.Lace.5676, adult female, vicinity of Makminasa, Pasimasunggu district, Kepulauan Selayar regency, Sulawesi Selatan province, Tanahjampea Island, Indonesia (07°05’20.3” S, 120°41’05.4” E, 103 m above sea level), collected by J. A. McGuire on 16 October 2005, 10:10 pm; MVZ 267742, MVZ 267744 (Fig.3), adult females, vicinity of Makminasa, Pasimasunggu district, Kepulauan Selayar regency, Sulawesi Selatan province, Tanahjampea Island, Indonesia (07°05’04.9” S, 120°40’58.3” E, 195 m above sea level), collected by Christopher J. Hayden on October 2005, 11:00 pm; MVZ 267749–50, adult males, Telkom tower road, Pasimasunggu district, Kepulauan Selayar regency, Sulawesi Selatan province, Tanahjampea Island, Indonesia (07°05’09.4” S, 120°39’29.2” E, 316 m above sea level), collected by J. A. McGuire, Christopher J. Hayden, and Ted Townsend on 18 October 2005, 8:30 pm; MVZ 267752, adult female, Telkom tower road, Pasimasunggu district, Kepulauan Selayar regency, Sulawesi Selatan province, Indonesia (07°05’12.3” S, 120°39’11.1” E, 251 m above sea level), collected by J. A. McGuire, Christopher J. Hayden, and Ted Townsend on 18 October 2005, 9:07 pm. 
DiagnosisDiagnosis. Cyrtodactylus tanahjampea sp. nov. is a medium size species reaching 76.1 mm and can be readily distinguished from the congeners on Sulawesi, the Moluccas, and the Lesser Sunda Islands by the following unique combination of characters: (1) brachium and antebrachium tuberculated, (2) ventrolateral folds with tubercles, (3) 20–23 irregularly aligned rows of keeled tubercles, (4) 31–34 paravertebral tubercles, (5) 29–34 ventral scales between ventrolateral folds, (6) no precloacal depression, (7) enlarged precloacofemoral scales in a continuous series, (8) males with 20–24 precloacofemoral pores in wide Ʌ-shape, (9) enlarged post precloacal scales present, (10) 19–21 fourth toe subdigital lamellae, (11) absence of enlarged transversely median subcaudals, (12) tail not prehensile, (13) tubercles extend over anterior about 71 % of original tail’s length, and (14) the original tails reach 147 % of SVL.

Species comparisons. Cyrtodactylus tanahjampea sp. nov. can be readily distinguished from all recognized congeners occurring on Sulawesi, the Moluccas, and the Lesser Sunda Islands (detailed diagnostic characters for the species from these regions are presented in Table 2) except for C. halmahericus (Mertens, 1929) from which it differs by possessing EPFS, PP*, FP*, and in lacking enlarged median subcaudal scales on original tails (characters that are occur on males only are donated by *). The new species is easily differentiated from C. halmahericus by having strongly keeled dorsal tubercles (vs. keeled tubercles), fewer EPFS (40–45 vs. 48–53), precloacofemoral pores* (28–34 vs. 48–53) and in lacking a precloacal depression as opposed to having a deep precloacal groove. Further, it differs from C. deveti in its much smaller body size (76.1 vs. 106 mm SVL) and in lacking enlarged transverse median subcaudal scales (as opposed to presence of enlarged transverse median subcaudal scales). The new species is also easily distinguished from C. nuaulu by having EPFS as opposed to presence of EPS, PFP* as opposed to presence of PP*, and in males lacking a precloacal depression as opposed to having a deep precloacal groove. Among the recognized species occurring in the Sulawesi region, Cyrtodactylus tanahjampea is similar to C. fumosus (data from Mecke et al. 2016b) in body size (< 80 mm SVL), in having of precloacofemoral pores in a continuous series, and in having a blotched dorsal pattern, but differs from that species in the presence of tubercles in the ventrolateral folds, in lacking a precloacal depression, in having more DorsT (20–23 vs. 4–7), and in having fewer VentS (28–34 vs. 37–50). Further, these species are also readily differentiated in the arrangement of their chin shields, with C. tanahjampea having paired elongated primary postmentals that contact medially for 50% of their posterior sections as opposed to having ~70% contact posteromedially in C. fumosus. The new species differs from C. batik (data from Iskandar et al. 2011; reexamine specimens see appendix 1) in being much smaller (76.1 vs. 114.6 mm), in having a blotched as opposed to a banded dorsal color pattern, in having fewer VentS (28–34 vs. 48–57), by the presence of EPFS (vs. lack of EPFS), PFP* (vs. poreless), and in lack enlarged transversely median subcaudal scales (as opposed to presence of enlarged transversely median subcaudals). The new species differs from C. hitchi in having tubercles on the dorsal surface of the brachium as opposed to the brachium untuberculated, EPFS as opposed to presence of EPS, PFP* as opposed to poreless both on precloacal and femoral, blotched as opposed to banded, and in lacking enlarged transverse median subcaudals as opposed to transversely median subcaudals arranged in a single row. Cyrtodactylus tanahjampea is further differentiated from C. jellesmae by having pores, EPFS, fewer VentS (28–34 vs. 40–50), and a blotched as opposed to striped dorsal pattern. It differs from C. tahuna in having EPFS and PFP* as opposed to a discontinuous series of EPS–EFS and a discontinuous series of PP*–FP*. From C. spinosus, the new species can be distinguished by its smaller body size (76.1 vs. 83.2 mm), its lack of spine-like tubercles scattered over the dorsum, in having PFP* and in lacking a precloacal depression. The new species differs from C. wallacei by its much smaller size (76.1 vs. 114 mm), by having PFP* as opposed to lacking pores, and in lacking enlarged transversely median subcaudal scales. The new species differs from all recognized species occuring in the Lesser Sundas in having a continuous series of precloacal and femoral pores in males. Cyrtodactylus tanahjampea differs from all species from this region except C. darmandvillei by having tubercles on the dorsal sufaces of the brachium. The new species can easily be distinguished from C. celatus (data from Kathriner et al. 2014, Rösler & Kaiser 2016, and Mecke et al. 2016a) by its much larger body size (76.1 vs. 38–44 mm), in having EPFS, in the absence of a precloacal depression as opposed to presence of a precloacal groove in males, and in having more lamellae under the fourth toe (18–23 vs. 15–18). From C. darmandvillei (data from Mecke et al.2016), the new species differs by lacking enlarged transverse median subcaudals as opposed to having enlarged transverse median subcaudals, and in having pores in males as opposed to being poreless in both sexes. From C. gordongekkoi (data from Das 1993, Biswas 2007, and Mecke et al. 2016a), the new species differs by the presence of tubercles on the dorsal surface of the brachium as opposed to absence and having precloacofemoral pores in males as opposed to being poreless in both sexes. From C. laevigatus (data from Kathriner et al. 2014 and Mecke et al. 2016a), the new species differs in its much larger body size (76.1 vs. 38.5 mm) and in having precloacofemoral pores in males as opposed to no pores in either sex. From C. tambora, the new species differs in its larger body size (76.1 vs. 39.4 mm), having precloacofemoral pores in males as opposed to presence of only precloacal pores in males, and in lacking a precloacal depression as opposed to presence of a precloacal groove in males. From C. wetariensis (Dunn, 1927) (data from Mecke et al. 2016a), the new species differs by having the brachium tuberculated as opposed to lacking tubercles on the dorsal surface of the brachium and by having precloacal pores and femoral pores arranged in a continuous series in males as opposed to being discontinuously arranged.
We also compare the new species to the other congeners occuring west and east of the Wallacea region. Enlarged precloacofemoral scales are present in Cyrtodactylus tanahjampea, whereas C. boreoclivus Oliver, Mumpuni, Krey & Richards, 2011, C. consobrinus (Peters, 1871), C. elok Dring, 1979, C. ingeri Hikida, 1990, C. majulah Grismer, Wood & Lim, 2012, C. matsuii Hikida, 1990, C. pubisulcus Inger, 1958, C. quadrivirgatus Taylor, 1962, C. rosichonariefi Riyanto, Grismer & Wood, 2015, C. semiadii Riyanto, Bauer & Yudha, 2014, C. sermowaiensis (De Rooij, 1915), and C. yoshii Hikida, 1990 lack such a series. The subcaudal scales are not transversely enlarged in C. tanahjampea, a condition that is shared with C. klakahensis Hartmann, Mecke, Kieckbusch, Mader & Kaiser, 2016, C. loriae (Boulenger, 1896), C. marmoratus Gray, 1831, C. novaeguineae (Schlegel, 1837), C. petani Riyanto, Grismer & Wood, 2015, C. psarops Harvey, O’Connell, Barraza, Riyanto, Kurniawan & Smith, 2015, C. rosichonariefei Riyanto, Grismer & Wood, 2015, C. semiadii, and C. semicinctus Harvey, O’Connell, Barraza, Riyanto, Kurniawan & Smith, 2015. In contrast, transversely enlarged subcaudals are present in C. boreoclivus, C. brevipalmatus (Smith, 1923), C. consobrinus, C. hikidai Riyanto, 2012, C. ingeri Hikida, 1990, C. malayanus (de Rooij, 1915), and C. rex Oliver, Richards, Mumpuni & Rösler, 2016. 
Comment 
EtymologyThe specific name tanahjampea refers to Tanahjampea Island, the only locality at which this species is known to occur. 
References
  • RIYANTO, AWAL; AMIR HAMIDY, JIMMY A. MCGUIRE 2018. A New Bent-toed Gecko (Cyrtodactylus: Squamata: Gekkonidae) from the Island of Tanahjampea, South Sulawesi, Indonesia. Zootaxa 4442 (1): 122–136 - get paper here
 
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