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Cyrtodactylus tripuraensis AGARWAL, MAHONY, GIRI, CHAITANYA & BAUER, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesTripura bent-toed gecko 
SynonymCyrtodactylus tripuraensis AGARWAL, MAHONY, GIRI, CHAITANYA & BAUER 2018
Cyrtodactylus sp. Tripura — AGARWAL et al. 2014 
DistributionIndia (Tripura), Bangladesh

Type locality: Chobimura (23.55039°N, 91.62054°E; ca. 36 m asl.), Gomati district, Tripura state, India  
Reproduction 
TypesHolotype: BNHS 2244, adult male, collected by A. Datta-Roy, T. Khichi, NPI Das and I. Agarwal, 10 November 2010. Paratypes. All from Tripura state, India, same collectors as holotype: BNHS 2230, adult female, from Rowa Wildlife Sanctuary (24.29062°N 92.16496°E; ca. 56 m asl.), North Tripura district, 02 October 2010; BNHS 2236– 2238, BNHS 2242, and BNHS 2243, five adult males, BNHS 2239–2241, three adult females, Sepahijala Wildlife Sanctuary (23.65906°N 91.33167°E; ca. 29 m asl.), Sepahijala district, 06–08 November 2010; BNHS 2245, adult male, Gumti (23.42661°N 91.81937°E; ca. 40 m asl.), Gomati district, 11 November 2010. 
DiagnosisDiagnosis and comparison against regional congeners. A moderate sized Cyrtodactylus, snout-vent length to 70.7 mm; body and limbs relatively slender, digits short; one pair of enlarged postmental scales in broad contact behind mental; 19–21 rows of circular tubercles at mid-dorsum, bluntly conical and weakly keeled throughout; 44– 54 paravertebral tubercles; 35–43 mid-body ventral scale rows; no precloacal groove; males with 29–37 precloacofemoral pores in a single series, females with 19–29 pitted/pored scales. Four to nine basal and 11–16 apical subdigital lamellae beneath Digit IV of pes with one or two intervening rows of non-lamellar granular scales between basal and distal lamellae series. Subcaudals larger than dorsal tail scales, not in an enlarged median series (on original tail). Dorsal pattern of 7–10 paired transversely arranged dark spots that may be irregular and broken up, occasionally alternating with lighter spots. Original tail with 11–13 alternating dark and light bands, last few light bands almost white.
Cyrtodactylus tripuraensis sp. nov. differs from the following species by the presence of a continuous series of precloacofemoral pores (vs. precloacal pores separated from femoral pores by a diastema of multiple poreless scales): C. annandalei, Cyrtodactylus bhupathyi sp. nov., C. fasciolatum, C. gubernatoris, C. russelli, and C. slowinskii. The large number of precloacofemoral pores on males of the new species (29–37) separates it from C. brevidactylus (8 PcP), C. cayuensis (6–9 PcP), Cyrtodactylus himalayicus comb. nov. (10 PcP), C. khasiensis (10– 12 PcP), C. mandalayensis (8 PcP + single enlarged pore-bearing scale at the posterior apex of PcP), C. markuscombaii (7 PcP), C. martinstolli (0–8 PcP), and C. wakeorum (12 PcP). Cyrtodactylus tripuraensis sp. nov. differs from C. tamaiensis by having slightly fewer precloacofemoral pores (29–37 vs. 40), male body size (SVL 55.4–65.0 mm vs. 90.0 mm), longer trunk (TrL/SVL mean 0.49, range 0.46–0.52 vs. 0.41) and dorsal pattern (8–10 irregular paired dark spots vs. irregular dark and light reticulations), and from C. gansi in the absence of a precloacal groove (vs. presence of precloacal groove on males). Cyrtodactylus tripuraensis sp. nov. is most similar to C. ayeyarwadyensis from which it differs by a slightly higher number of precloacofemoral pores on males (29– 37 vs. 10–28), the presence of pores in females (vs. absence of pores), and fewer dorsal tubercle rows (19–21 vs. 22–24). Cyrtodactylus tripuraensis sp. nov. differs from C. ayeyarwadyensis sensu lato (Mahony et al. 2009a) by the presence of precloacofemoral pores on females (vs. absence of pores). 
CommentHabitat: <1.5 m from the ground and mainly from muddy road cuts in secondary and degraded moist deciduous forest; the individual from Rowa was on the trunk of a coppiced tree and one from Gumti was found under a rock in a hill forest approximately 10 m from the road. 
EtymologyThe species epithet is a toponym for Tripura, the Indian state in which all currently known populations are found. 
References
  • Agarwal, Ishan; Aaron M. Bauer, Todd R. Jackman, K. Praveen Karanth 2014. Insights into Himalayan biogeography from geckos: A molecular phylogeny of Cyrtodactylus (Squamata: Gekkonidae). Molecular Phylogenetics and Evolution - get paper here
  • AGARWAL, ISHAN; STEPHEN MAHONY, VARAD B. GIRI, R. CHAITANYA, AARON M. BAUER 2018. Two new species of bent toed geckos, Cyrtodactylus Gray, 1827 (Squamata: Gekkonidae) from Northeast India with comments on name-bearing types from the region. Zootaxa 4420 (3): 334-356 - get paper here
  • Al-Razi H, Maria M, Hasan S 2018. First record of the recently described Cyrtodactylus tripuraensis Agarwal, Mahony, Giri, Chaitanya & Bauer, 2018 (Squamata, Gekkonidae) in Bangladesh. Check List 14(6): 1105-1108 - get paper here
 
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