Delma butleri STORR, 1987

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Higher Taxa	Pygopodidae, Gekkota, Sauria, Squamata (lizards)
Subspecies
Common Names	E: Unbanded Delma, Butler's Legless Lizard, Spinifex Snake-lizard
Synonym	Delma butleri STORR 1987: 346 Delma haroldi STORR 1987: 350 Delma butleri — KLUGE 1993 Delma butleri — COGGER 2000: 286 Delma haroldi — MARYAN et al. 2007 Delma haroldi — WILSON & SWAN 2010: 130 Delma butleri — WILSON & SWAN 2010
Distribution	Australia (New South Wales, Northern Territory, S Queensland, South Australia, Victoria, Western Australia) Type locality: 5 km SE Gnaraloo, WA, in 23°52'S 113°32'E. haroldi: Australia (SW Northern Territory, Western Australia);Type locality: 1 km NNW Daroo, WA, in 22°41'S, 115°20'E.
Reproduction	oviparous
Types	Holotype: WAM R76742 Holotype: WAM R85094 [haroldi]
Diagnosis	Diagnosis: A small Delma without dark or pale bands across top of head and neck. Distinguishable from D. nasuta by fewer and differently patterned loreals, shorter and darker snout, dark brown upper labials variably marked with white (rather than brownish white, narrowly edged above with dark brown), dorsals dark brown finely edged with blackish (rather than pale brown, spotted dark brown) and ventrals and subcaudals never dark-edged. Distinguishable from D. grayii G.M. Storr by finely dark-edged dorsals and little or no development of ventrolateral vertical barring behind ear. Additional details (593 characters) are available for collaborators and contributors. Please contact us for details.
Comment	Limb morphology: Limbless. Synonymy: Shea (1991) considered D. haroldi a junior synonym of D. butleri. which was confirmed by Brennan et al. 2016), also it has been considered as valid by various recent authors. The range of D. haroldi lies within the range of D. butleri. NCBI taxonID: 207581 [haroldi]
Etymology	After naturalist W.H. (Harry) Butler, who was first to collect this species for the Western Australian Museum (on 30 December 1961 at Queen Victoria Spring). Delma haroldi was named after naturalist Gregory Harold, co-collector of the holotype.
References	Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here Brennan, Ian G.; Aaron M. Bauer, Todd R. Jackman 2015. Mitochondrial introgression via ancient hybridization, and systematics of the Australian endemic pygopodid gecko genus Delma. Molecular Phylogenetics and Evolution 94 (2016) 577–590 - get paper here Cogger, H. G. 2014. Reptiles and Amphibians of Australia, 7th ed. CSIRO Publishing, xxx + 1033 pp. - get paper here Cogger, H.G. 2000. Reptiles and Amphibians of Australia, 6th ed. Ralph Curtis Publishing, Sanibel Island, 808 pp. Dittmer, D. E., Chapman, T. L. & Bidwell, J. R. 2020. In the shadow of an iconic inselberg: Uluru’s shadow influences climates and reptile assemblage structure at its base. J. Arid Environ. 181, 104179 (2020) - get paper here Ellis, Ryan J.; Paul Doughty and Aaron M. Bauer 2018. An annotated type catalogue of the geckos and pygopods (Squamata: Gekkota: Carphodactylidae, Diplodactylidae, Gekkonidae, Pygopodidae) in the collection of the Western Australian Museum. Records of the Western Australian Museum 33: 051–094 - get paper here Jennings, W.B.; Pianka, E.R. & Donnellan, S. 2003. Systematics of the lizard family Pygopodidae with implications for the diversification of Australian temperate biotas. Systematic Biology 52: 757-780 - get paper here Maryan, B. 2005. A HERPETOFAUNA HOTSPOT, THE CENTRAL WEST COAST OF WESTERN AUSTRALIA. Western Australian Naturalist 25 (1): 1-24 - get paper here Maryan,B., Aplin,K., & Adams,M. 2007. Two new species of the Delma tincta group (Squamata: Pygopodidae) from northwestern Australia. Rec. West. Austr. Mus. 23: 273-305 - get paper here Pianka, E. R. 1969. Habitat specificity, speciation, and species density in Australian desert lizards. Ecology 50 (3): 498-502 - get paper here Pianka, E. R. 1986. Ecology and Natural History of Desert Lizards. Analyses of the Ecological Niche and Community Structure. Princeton University Press, Princeton, New Jersey - get paper here Pianka, E. R. 2010. Notes on the ecology of four species of pygopodid lizards in the Great Victoria Desert. Western Australian Naturalist 27: 175-181 - get paper here Shea, G. M. 1991. Revisionary notes on the genus Delma (Squamata: Pygopodidae) in South Australia and the Northern Territory. Rec. South Austral. Mus. 25: 71-90 - get paper here Storr, G. M. 1987. Three new legless lizards (Pygopodidae) from Western Australia. Rec. West. Austr. Mus. 13 (4): 345—355. - get paper here Swan, G.; Sadlier, R.; Shea, G. 2017. A field guide to reptiles of New South Wales. Reed New Holland, 328 pp. Wilson, S. & Swan, G. 2010. A complete guide to reptiles of Australia, 3rd ed. Chatswood: New Holland, 558 pp.
External links	IUCN Red List National Center for Biotechnology Information http://www.amonline.net.au/herpetology/research/pdf/pygopodidae.pdf http://www.kingsnake.com/oz/lizards/legless/legless.htm http://www.museum.vic.gov.au/bioinformatics/lizards/images/butllive1.htm http://www.museum.vic.gov.au/bioinformatics/lizards/images/butllive2.htm http://www.arod.com.au/arod/reptilia/Squamata/Pygopodidae/Delma/butleri Google images

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