Dendragama dioidema HARVEY, SHANEY, SIDIK, KURNIAWAN & SMITH, 2017
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Dendragama dioidema
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| Higher Taxa | Agamidae (Draconinae), Sauria, Iguania, Squamata (lizards) |
| Subspecies | |
| Common Names | |
| Synonym | Dendragama dioidema HARVEY, SHANEY, SIDIK, KURNIAWAN & SMITH 2017 |
| Distribution | Indonesia (Sumatra) Type locality: Bukit Sama, Kampung Telege Atu, Kebayakan, Kabupaten Aceh Tengah, Provinsi Aceh, Indonesia, 4.66512°N, 96.80937°E, 1567 m |
| Reproduction | oviparous; Harvey et al. 2017 concluded that Dendragama australis normally has a clutch size of four and reaches sexual maturity around 60 mm SVL. Presence of eggs in pairs and in up to four developmental stages strongly suggests that female D. australis produce multiple clutches per year. |
| Types | Holotype: MZB 13814, adult male (collector’s tag ENS 19450; Fig. 12), collected 5 August 2015 by I. Sidik, I. Fonna, and P. Thammachoti. Paratypes: (42) (note missing ° symbols in coordinates); Three males (MZB 13794; UTA 63448, 63449), six females (MZB 13804, 13805; UTA 63438, 63440, 63441, 63446), and 15 unsexed specimens (MZB 13795– 13803; UTA 63439, 63442–63445, 63447) from Kute Baru, Linge, along road from Takengon to Isaq, Kabupaten Aceh Tengah, Provinsi Aceh, Indonesia, 4.528758N, 96.853168E, 1827 m, collected on 5 August 2015 by E. Wostl, I. Fonna and M. Ikhsan; one unsexed specimen (UTA 63450) from foot of Berni Terlong, near Desa Rambune, Kabupaten Bener Meriah, Provinsi Aceh, Indonesia, 4.76488N, 96.801968E, 1471 m, collected on 7 August 2015 by G.C. Sarker, I. Sidik, I. Fonna and M. Ikhsan; four male (MZB 13807, 13808, 13810; UTA 63452) and two unsexed specimens (MZB 13809; UTA 63453) from Beutong Ule, high point on Meulaboh-Takengon road, Kabupaten Nagan Raya, Provinsi Aceh, Indonesia, 4.383678N, 96.516338E, 1950 m, collected on 5 August 2015 by E. Wostl, E.N. Smith, G.C. Sarker and P. Thammachoti; one male (MZB 13811), three females (UTA 63456–63458), and three unsexed specimens (MZB 13812, 13813; UTA 63455) from Bukit Sama, Kampung Telege Atu, Kebayakan, Kabupaten Aceh Tengah, Provinsi Aceh, Indonesia, 4.66512–4.665838N, 96.80627–96.809378E, collected 5 August 2015 by I. Sidik, I. Fonna, and P. Thammachoti; one male (UTA 63460), one female (MZB 13815), and two unsexed specimens (MZB 13816; UTA 63459) from Hutan Timang Gajah, Gunung Burni Telong, Kabupaten Bener Meriah, Provinsi Aceh, Indonesia, 4.77122–4.771428N, 96.80907–96.810178E, 1875–1957 m, collected on 7 August 2015 by E. Wostl, P. Thammachoti, A.M. Khadafi, and I. Fonna. |
| Diagnosis | Diagnosis: The following combination of characters distinguishes Dendragama dioidema from its congeners: (1) enlarged scales below eye broadly contacting supralabials; (2) tympanum relatively large (11–16% of head length) and close to eye (orbit–meatus 23–29% of head length); (3) sublabial tubercular scales 0–2, usually 1, in front of postrictal modified scale; (4) gulars large, 16–22; (5) dorsal crest of 9–19 projecting scales, serrate with obvious gaps between enlarged projecting scales; (6) dorsolateral crest of more or less continuous posttympanic and dorsolateral series; (7) scales around midbody 57–77; (8) scales of lower flanks heterogenous in size, smooth to feebly keeled; (9) ventrals smooth or very feebly keeled, 49–63; (10) no white sublabial stripe; however, one or two white or pale yellow spots (corresponding to sublabial tubercular scale and postrictal modified scales) usually present in area below rictus and ear; (11) no distinctive prescapular blotch edged posteriorly in white; band crossing neck interrupted by narrow white or yellow line; (12) proximal half of tail with 6– 10 dark brown or green bands; (13) buccal epithelium cream, tongue pink to red. (Harvey et al. 2017) |
| Comment | Distribution: see map in Shaney et al 2020: 130 (Fig. 1). |
| Etymology | The new name dioidema is a masculine noun in apposition derived from the Greek prefix di, meaning two, and noun oidema, meaning protuberance. The name refers to the pair of distinctive tubercular scales positioned slightly below and behind the rictus in this species. Unlike some congeners, Dendragama dioidema usually has a single, large sublabial tubercular scale positioned in front of the postrictal modified scale. |
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