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Diploderma ngoclinensis (ANANJEVA, ORLOV & NGUYEN, 2017)

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Higher TaxaAgamidae (Draconinae), Sauria, Iguania, Squamata (lizards) 
Subspecies 
Common Names 
SynonymJapalura ngoclinensis ANANJEVA, ORLOV & NGUYEN 2017
Diploderma ngoclinensis — WANG et al. 2018 
DistributionVietnam (Kon Tum Province)

Type locality: above Mang Xang Village, Ngoc Linh Mountain, Dac Glei District, Kon Tum Province, Vietnam (15°05' N 107°57' E, elevation 1650 m),  
Reproductionoviparous 
TypesHolotype: ZISP 26230 (adult female), collected on 25 March 2004 by Alexei V. Abramov (Figures 2 and 5). Paratypes ZISP 26231 (adult female) and VNMH 3110 (subadult female) from above Mang Xang Village, Ngoc Linh Mountain, Dac Glei District, Kon Tum Province, Vietnam (15°05' N 107°57' E, elevation 1650 m), on 25 March 2004 by Alexei V. Abramov (Figures 3–5). 
DiagnosisDiagnosis Medium-sized, long-tailed Japalura species with relatively long limbs, that differs from all other species of this genus by the following combination of characters: tympanum hidden, transverse gular fold absent, enlarged subocular scale row present. Third and fourth fingers have equal length. Very low dorsal crest from 5–6 slightly enlarged serrated scales.

Comparison Japalura ngoclinensis has tympanum hidden what differs it from following species: J. dasi Shah et Kästle, 2002), J. major (Jerdon, 1870), J. tricarinata (Blyth, 1853), J. kumaonensis (Annandale, 1907), J. varcoae (Boulenger, 1918) and J. dymondi (Boulenger, 1906), possessing exposed tympanum (Smith, 1935; Manthey et al., 2012). Additionally from: J. dasi, J. kumaonensis and J. varcoae by the presence of a transverse gular fold (vs. absent); from J. major by a fewer T4S ≤ 22 (vs. ≥ 23); from J. tricarinata by shorter limbs FLL/SVL ≤ 45.5% (vs. ≥ 50%) and HLL/SVL ≤ 82% (vs. ≥ 86%) and from J. major by a longer tail length TL/SVL ≥ 226% (vs. ≤ 182%) (Smith, 1935; Mahony, 2009, 2010; Manthey et al., 2012).
According to detailed data arrangement from Manthey et al., 2012 we made particular comparison within the group of species with tympanum concealed and absence or presence of transverse gular fold:
From the species with tympanum hidden and presence of transverse gular fold J. batangensis Li, Deng, Wu et Wang, 2001, J. brevicauda Manthey, Denzer, Hou et Wang, 2012, J. flaviceps Barbour et Dunn, 1919, J. hamptoni Smith, 1935, J. yulongensis Manthey, Denzer, Hou et Wang, 2012, J. splendida Barbour et Dunn, 1919, J. zhaoermii Goa et Hou, 2002, J. vela Wang, Jiang et Che, 2015, J. laeviventris Wang, Jiang, Siler, et Che, 2016 and J. iadina Wang, Jiang, Siler et Che, 2016; it differs by the absence of such a fold.
Additionally from J. batangensis by a narrower head HW≤18%(vs.≥21%),alessernumberofIL≤9(vs.≥ 10) and by a higher number of MD ≥ 54 (vs. ≤ 44); from J. brevicauda J. ngoclinensis differs by the possessing longer tail ≥ 226% (vs. ≤ 145%); from J. flaviceps by a narrower head HW/SVL ≤ 18% (vs. ≥ 19%), from J. hamptoni by a higher number of MD ≥ 54 (vs.44) and fewer F4S ≥ 20 (vs. 24); from J. zhaoermii by a narrower head HW/SVL ≤ 18% (vs. ≥ 20%), higher number of MD ≥54(vs.≤47)andfewerF4S≥20(vs.≥24); fromJ. yulongensis by a higher number of MD ≥ 54 (vs. 35–42); from J. vela by more narrow head HW/SVL ≤ 17% (vs. ≥ 21%), shorter forelimbs ≤ 45% (vs. ≥ 45%) and a higher number of MD ≥ 54 (vs. ≤ 51); from J. laeviventris by lesser number of MD ≤ 56 (vs. ≥ 57) and from J. iadina by a shorter relative snout length SEL/HL ≥ 41% (vs. 34.9%–40.2%).
Comparison within the group of species with tympanum concealed and absence of transverse gular fold: Japalura ngoclinensis differs
from J. andersoniana Annandale, 1905 by absence of arrangement of dorsal pholidosis with v-shaped rows of enlarged dorsals (vs. enlarged dorsal scales in v-shaped rows) and shorter hindlimbs HLL/SLV ≤ 82% (vs. ≥ 88%); from J. chapaensis Bourret, 1937 by fewer T4S ≤ 26 (vs. ≥ 28);
from J. fasciata Mertens, 1926, by slightly smaller adult size of females, SVL to 58.7 mm (vs. to 71 mm), higher number of MD ≥ 54 (vs. ≤ 38), and absence of clear white transverse band (vs. presence of such a band);
from J. grahami Steineger 1924 by more NSL ≥ 1 (vs. absent) and MD present (vs.absent); from J. micangshanensis Song 1987 by narrower head width HW/SVL ≤ 18% (vs. ≥ 21%);
from J. otai Mahony, 2009 by narrower head width HW/SVL ≤ 18% (vs. ≥ 21%);
from J. planidorsata Jerdon, 1870 by a larger SVL ≥ 57 mm (vs. ≤ 53 mm);
from J. sagittifera Smith, 1940 by the absence of a light stripe from the eye to the angle of the mouth (vs. present) and a little fewer MD ≤ 52 (vs. ≥ 53);
from J. variegata Gray, 1853 by a little more ≥ 54 (vs. ≤ 54) mm; from J. yunnanensis (Anderson, 1878) by presence of NSL≥1(vs.0)byahighernumberofMD≥54(vs.≤ 43).
From the East Asian insular species, it differs
from J. brevipes Gressit, 1936, J. luei Ota, Cheng et Shang, 1998 and J. makii Ota, 1989 by having more NSL ≥ 1 (vs.0);
from J. polygonata Hallowell, 1861 and J.swinhonis Günther, 1864 by absence (vs. presence) of a light lateral stripe and more clearly pronounced nuchal crest and by shorter forelimbs ≤ 45% (vs. ≥ 47.6% and ≥ 45% respectively).
The character of enlarged row of scales offered by Mahony (2010) as one of three scale characters which in different combinations can be used in attempt to stabilise the generic status of some systematically problematic draconin species. It is present in J. ngoclinensis as well as in the following East Asian Japalura s.l. species with this enlarged row of scales: J. brevipes, J. chapaensis, J. fasciata, J. hamptoni, J. luei, J. makii, J. splendida and J. yunnanensis. 
CommentStatus unclear. May be a synonym of D. splendidum (Wang et al. 2018).

Abundance: only known from its original description (Meiri et al. 2017). 
EtymologyThe specific epithet refers to the distribution of the new species in the Ngoc Linh Mountain, Kon Tum Province, Vietnam. 
References
  • ANANJEVA, Natalia B.; Nikolay L. ORLOV and Tao Thien NGUYEN 2017. A New Species of Japalura (Agamidae: Lacertilia: Reptilia) from Central Highland, Vietnam Asian Herpetological Research 8 (1): 14–21 - get paper here
  • DENZER, WOLFGANG; ULRICH MANTHEY, PATRICK D CAMPBELL 2019. Catalogue of type specimens of the agamid lizard genus Japalura s. l. (Squamata: Agamidae: Draconinae). Zootaxa 4612 (1): 109-125 - get paper here
  • Meiri, Shai; Aaron M. Bauer, Allen Allison, Fernando Castro-Herrera, Laurent Chirio, Guarino Colli, Indraneil Das, Tiffany M. Doan, Frank Glaw, Lee L. Grismer, Marinus Hoogmoed, Fred Kraus, Matthew LeBreton, Danny Meirte, Zoltán T. Nagy, Cristiano d 2017. Extinct, obscure or imaginary: the lizard species with the smallest ranges. Diversity and Distributions - get paper here
  • Wang, Kai; Jing Che, Simin Lin, V Deepak, Datta-Roy Aniruddha, Ke Jiang, Jieqiong Jin, Hongman Chen, Cameron D Siler; 2018. Multilocus phylogeny and revised classification for mountain dragons of the genus Japalura s.l. (Reptilia: Agamidae: Draconinae) from Asia. Zoological Journal of the Linnean Society, , zly034, - get paper here
 
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