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Dipsas copei (GÜNTHER, 1872)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common Names 
SynonymLeptognathus Copei GÜNTHER 1872: 30
Dipsas copei — PETERS 1960: 58
Dipsas copii — ROZE 1966: 11
Dipsas copei — PETERS & OREJAS-MIRANDA 1970: 86
Dipsas copei — STARACE 1998: 167
Dipsas copei — KORNACKER 1999: 78
Dipsas copei — CLAESSEN 2003
Dipsas copei — HARVEY et al. 2008
Dipsas copei — WALLACH et al. 2014: 231
Dipsas copei — NOGUEIRA et al. 2019 
DistributionGuyana, Suriname, French Guiana, S Venezuela

Type locality: “Suriname ?” (fide KORNACKER 1999).  
Reproductionoviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: BMNH 1946.1.21.4 formerly 66.8.14.329), male, from collection of Dr. van Lidth de Jeude. 
Diagnosis 
CommentLeptognathus alternans FISCHER 1885 has been removed from the synonymy of D. incerta by PASSOS et al. (2004). D. copei has been synonymized with D. incerta by PASSOS et al. 2004 and revalidated by HARVEY 2008. However, D. copei may be a synonym of D. praeornata (here still a synonym of Dipsas latifrontalis).The neotype designated by PASSOS et al. (2004) is invalid fide HARVEY 2008.

D. copei is not listed by GASC & RODRIGUES 1980.

Habitat: fully arboreal (Harrington et al. 2018). 
EtymologyNamed after Edward Drinker Cope (1840-1897), American herpetologist. See Osborn 1929 and Vetter 2003 for biographical notes. 
References
  • Beolens, Bo; Michael Watkins, and Michael Grayson 2011. The Eponym Dictionary of Reptiles. Johns Hopkins University Press, Baltimore, USA - get paper here
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • Claessen, H. 2003. Slangen van de Guyana’s. Deel V. Lacerta 61 (4): 123-133 - get paper here
  • Cole, Charles J.; Carol R. Townsend, Robert P. Reynolds, Ross D. MacCulloch, and Amy Lathrop 2013. Amphibians and reptiles of Guyana, South America: illustrated keys, annotated species accounts, and a biogeographic synopsis. Proceedings of the Biological Society of Washington 125 (4): 317-578; plates: 580-620 - get paper here
  • Fischer, J. G. 1885. Ichthyologische und herpetologische Bemerkungen. V. Herpetologische Bemerkungen. Jahrb. Hamburg. Wiss. Anst. 2: 82-121 - get paper here
  • Günther,A. 1872. Seventh account of new species of snakes in the collection of the British Museum. Ann. Mag. Nat. Hist. (4) 9: 13-37 - get paper here
  • Harrington, Sean M; Jordyn M de Haan, Lindsey Shapiro, Sara Ruane 2018. Habits and characteristics of arboreal snakes worldwide: arboreality constrains body size but does not affect lineage diversification. Biological Journal of the Linnean Society 125 (1): 61–71 - get paper here
  • Harvey, Michael B. 2008. New and Poorly Known Dipsas (Serpentes: Colubridae) from Northern South America. Herpetologica 64 (4): 422-451 - get paper here
  • Harvey, Michael B. and Dirk Embert 2008. Review of Bolivian Dipsas (Serpentes: Colubridae), with Comments on Other South American Species. Herpetological Monographs 22 (1): 54-105 - get paper here
  • Harvey, Michael B.; Gilson Rivas Fuenmayor, José Rances Caicedo-Portilla, and José Vicente Rueda-Almonacid 2009. Systematics of the Enigmatic Dipsadine Snake Tropidodipsas perijanensis Alemán (Serpentes: Colubridae) and Review of Morphological Characters of Dipsadini. Herpetological Monographs 22 (1): 106-132 - get paper here
  • Jan, G. 1863. Elenco Sistematico degli Ofidi descriti e disegnati per l'Iconografia Generale. Milano, A. Lombardi. vii + 143 pp. - get paper here
  • Kornacker,P.M. 1999. Checklist and key to the snakes of Venezuela. PaKo-Verlag, Rheinbach, Germany, 270 pp.
  • Lancini,A.R. & Kornacker,P.M. 1989. Die Schlangen von Venezuela. Armitano Editores C.A., Caracas, 1-381
  • Natera-Mumaw, Marco; Luis Felipe Esqueda-González & Manuel Castelaín-Fernández 2015. Atlas Serpientes de Venezuela. Santiago de Chile, Dimacofi Negocios Avanzados S.A., 456 pp. - get paper here
  • Nogueira, Cristiano C.; Antonio J.S. Argôlo, Vanesa Arzamendia, Josué A. Azevedo, Fausto E. Barbo, Renato S. Bérnils, Bruna E. Bolochio, Marcio Borges-Martins, Marcela Brasil-Godinho, Henrique Braz, Marcus A. Buononato, Diego F. Cisneros-Heredia, 2019. Atlas of Brazilian snakes: verified point-locality maps to mitigate the Wallacean shortfall in a megadiverse snake fauna. South American J. Herp. 14 (Special Issue 1):1-274 - get paper here
  • Osborn, H. F. 1929. Biographical memoir of Edward Drinker Cope 1840-1897. National Academy of Sciences 13: 127-317 (1930?)
  • Parker, H. W. 1935. The frogs, lizards, and snakes of British Guiana. Proc. Zool. Soc. London 1935: 505-530 - get paper here
  • Passos, P., D. S. Fernandes and U. Caramaschi 2004. The taxonomic status of Leptognathus incertus Jan, 1863, with revalidation of Dipsas alternans (Fischer, 1885) (Colubridae: Dipsadinae). Amphibia-Reptilia 25 (4): 381-393 - get paper here
  • Peters , J. A. 1960. The snakes of the subfamily Dipsadinae. Misc. Publ. Mus. Zool., Univ. Michigan (114): 224 pp. - get paper here
  • RIVAS, GILSON A.; CÉSAR R. MOLINA, GABRIEL N. UGUETO, TITO R. BARROS, CÉSAR L. BAR- RIO-AMORÓS & PHILIPPE J. R. KOK 2012. Reptiles of Venezuela: an updated and commented checklist. Zootaxa 3211: 1–64 - get paper here
  • Starace, Fausto 1997. Contribution to the study of the snakes of French Guyana II. The presence of Dipsas copei (Gunther, 1872) in French Guyana. Litteratura Serpentium 17 (5): 97-101, 18 (4): 104 (erratum)
  • Starace, Fausto 1998. Guide des Serpents et Amphisbènes de Guyane. IBIS Rouge Editions, Guadeloupe, Guyane, 450 pp.
  • Starace, Fausto 2013. Guide des Serpents et Amphisbènes de Guyane. Ibis Rouge Editions, Matoury, Guyane, ISBN 978-2-84450-407-4 - get paper here
  • Vetter, H. 2003. Edward Drinker Cope (1840-1897). Sekretär 3 (2): 21-30 - get paper here
  • Vidal, Nicolas; de Massary, Jean-Christophe; Marty, Christian 1998. Nouvelles especes de serpents pour la Guyane francaise. Revue Française d'Aquariologie Herpétologie 25 (3-4): 131-134 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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