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Epictia wynni WALLACH, 2016

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Higher TaxaLeptotyphlopidae, Epictinae, Epictini, Typhlopoidea, Serpentes, Squamata (snakes) 
Subspecies 
Common Names 
SynonymEpictia wynni WALLACH 2016: 298
Leptotyphlops phenops — KLAUBER 1940: 151 (part)
Leptotyphlops phenops — DIXON et al., 1972: 229
Leptotyphlops phenops — SMITH & SMITH, 1976: S-B-130 (part), S-F-34
Leptotyphlops phenops — CARABIAS-LILLO et al., 1999: 118.
Leptotyphlops phenops phenops — SMITH & TAYLOR 1945: 24 (part)
Leptotyphlops phenops phenops — SMITH & SMITH 1976: S-B-130 (part)
Leptotyphlops phenops phenops — YESCAS-LAGUNA 2007: 149.
Leptotyphlops goudotii phenops — PETERS et al., 1970 and 1986: 170 (part)
Leptotyphlops goudotii phenops — HAHN 1980: 16 (part)
Leptotyphlops goudoti — SMITH & SMITH 1976: S-B-129–130 (part)
Leptotyphlops goudoti — HAHN 1979: 230 (part)
Leptotyphlops goudoti — MONTES DE OCA & PÉREZ-RAMOS, 1998: 90ff (map)
Leptotyphlops goudoti — CARABIAS-LILLO et al., 1999: 118
Leptotyphlops goudoti phenops — SMITH & SMITH 1976: S-B-130 (part)
Leptotyphlops goudotii — HEDGES 1996: 111 (part)
Leptotyphlops goudotii — KÖHLER 2001b: 12, fig. 11B (part)
Leptotyphlops goudotii — KÖHLER 2008: 183, map (part)
Leptotyphlops goudotii — TINOCO-NAVARRO 2005: 24ff
Leptotyphlops goudotii — CONABIO 2009: 17 (part)
Leptotyphlops goudotii — DIXON & LEMOS-ESPINAL 2010: 286, map 71
Leptotyphlops goudotii — HERNÁNDEZ-SALINAS & RAMÍREZ-BAUTISTA 2013: 10, fig. 5B.
Leptotyphlops a[lbifrons] phenops — VILLAVICENCIO 2001: 131
Leptotyphlops goudotti (sic) — HUITZIL-MENDOZA 2007: 91
Epictia goudotii — RAMÍREZ-BAUTISTA et al. 2010: 83, fig. 122, pl. 83
Epictia goudotii — RAMÍREZ-BAUTISTA 2014: 200–201, photo 223
Epictia goudotii — WILSON et al. 2013: 44 (part)
Epictia goudotii — LEMOS-ESPINAL & SMITH, 2015: 7
Epictia phenops — WALLACH et al. 2014: 278 (part) 
DistributionMexico (N Querétaro, NW Hidalgo)

Type locality: Río Ayutla, 30 km north of Jalpan de Serra, Querétaro, Mexico, 21°25'01"N, 99°36'15"W, elevation 575 m  
Reproduction 
TypesHolotype: TCWC 32899 (field no. RLB 137), a 163 mm (LOA) specimen collected by R. L. Beck on 7 June 1970.
Paratypes (15): MEXICO: QUERÉTARO: Arroyo Seco, ca. 2 mi S Conca, between Hacienda X-Conca and Río Santa Maria, collected by Carl S. Lieb on 6 June 1979, UTEP H-20217 (field no. CSL 5708); 2.5 km NW Hacienda Conca, collected by Carl S. Lieb on 30 December 1971, TCWC H-37722 (field no. CSL 1201); nr. Río Santa Maria, W of Hacienda Conca, collected by Carl S. Lieb on 1 January 1972, TCWC H-37723 (field no. CSL 1223); Hacienda Conca, collected by Robert A. Thomas on 6 January 1974, TCWC H-45539 (field no. RAT 4892); Hacienda Conca, collected by Robert A. Thomas on 7 January 1974, H-45542 (field no. RAT 4895), H-45544 (field no. RAT 4897); Hacienda Conca, collected by Robert A. Thomas on 8 January 1974, H-45547 (field no. RAT 4912); El Trapiche, collected by S. Hayes on 4 January 1974, TCWC H-45525 (field no. RAT 4852); El Trapiche, collected by Robert A. Thomas on 5 January 1974, TCWC H-45526 (field no. RAT 4860), H-45528 (field no. RAT 4864), H-45530 (field no. RAT 4866), and H-45532–35 (field no. RAT 4868–71). 
CommentSynonymy: after Wallach 2016: 98. See this paper also for references.

Diagnosis: (1) scale row formula = 14-14-14; (2) midtail scale rows = 10; (3) total length = 125–163 (x– = 147.2) mm; (4) total midodrsals = 242–260 (x– = 249.8); (5) subcaudals = 14–20 (x– = 16.3); (6) relative body pro- portion = 43–58 (x– = 47.8); (7) relative tail length = 4.4%–6.9% (x– = 5.7%); (8) relative tail width = 2.5–4.3 (x– = 3.3); (9) relative rostral width = 0.29–0.42 (x– = 0.36); (10) relative eye size = 0.36–0.44 (x– = 0.41); (10) rostral subtriangular with truncated apex, not or just reaching eye level; (11) supralabials 2, moderate anterior supralabial reaching mid-eye level; (12) frontal hexagonal, as wide than long; (13) supraoculars large, twice as broad as deep, posterior borders parallel to supranasal borders; (14) widest anteriormost vertebral scale 5th; (15) parietals deeper than occipitals, oriented transversely; (16) infralabials 3; (17) cloacal shield subtriangular; (18) head brown, with a small yellow spot on rostral; (19) dorsum with 7 dark brown to black stripes composed of contiguous strings of triangular- or diamond-shaped spots bordered by moderate to broad zigzag yellow stripes (= 7 dark stripes); (20) venter uniform brown; (21) midbody stripe formula (7 + 0) and middorsal pattern (3D + 4L); (22) tail with a pale terminal spot covering the 0–1 (x– = 0.4) dorsal scales and 0–8 (x– = 4.0) ventral scales (ventral/dorsal ratio 10.0); and (23) apical spine a small thin spike.

Habitat: This species inhabits agricultural and cultivated areas, tropical deciduous and evergreen forest, and pine-oak forest (Montes de Oca and Pérez-Ramos, 1998; Hernández-Salinas and Ramírez-Bautista, 2013), as well as thorn-scrub regions with abundant acacia with scattered cacti and non-thorny shrubs (Dixon et al., 1972). UTEP H-20217 was found under a stone in an open pasture.

Comparisons: see Wallach 2016: 302 for detailed comparisons. 
Etymology“This species is dedicated to Addison H. Wynn, Museum Specialist (Collections Manager), Division of Amphibians and Reptiles, Department of Vertebrate Zoology, National Museum of Natural History, Smithsonian Institution (USNM), and commemorates his 35 years of service to the museum. Addison is an excellent researcher who works diligently on scolecophidians, and always has assisted me as a source of specimens, data, and ideas.” 
References
  • Wallach, Van. 2016. Morphological review and taxonomic status of the Epictia phenops species group of Mesoamerica, with description of six new species and discussion of South American Epictia albifrons, E. goudotii, and E. tenella (Serpentes: Leptotyphlopidae: Epictinae Mesoamerican Herpetology 3 (2): 216-374 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
 
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