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Eutropis greeri BATUWITA, 2016

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Higher TaxaScincidae, Mabuyinae, Scincoidea, Sauria, Squamata (lizards) 
Subspecies 
Common Names 
SynonymEutropis greeri BATUWITA 2016
Eutropis floweri — SOMAWEERA & SOMAWEERA, 2009, in part
Eutropis madaraszi — SOMAWEERA & SOMAWEERA, 2009, in part
Eutropis macularia — DAS et al., 2008, in part 
DistributionSri Lanka (Galle)

Type locality: Kombala-Kottawa Forest Reserve, Galle, Sri Lanka, 06°06’N, 80°15’E, 60 m elevation  
Reproduction 
TypesHolotype: WHT 7000, male, 69.0 mm SVL, 24 October 2005, collected by S. Batuwita.
Paratypes. (All from Sri Lanka) WHT 6957, 6997, 7009, two males and one female, 51.0, 70.0, 51.0 mm SVL, respectively, same location data as holotype, 8 April 2003, collected by S. Batuwita; WHT 6961, female, 58.0 mm SVL, same location data as holotype, 20 April 2002, collected by S. Batuwita and I. Alagiyawadu; WHT 6965, 6992, female and male, 47.0, 51.0 mm SVL, same location data as holotype, 29 March 2001, collected by S. Batuwita and I. Alagiyawadu; WHT 6987, male, 56.0 mm SVL, Beraliya Forest Reserve, Pituwala near Elpitiya, 068150N, 808120E, 80 m, 8 June 2001, collected by S. Batuwita; NMSL RSK 5, female, 70.0 mm SVL, Dilkanda?, May 1927; WHT 6986, 6990, 6993, one female and two males, 55.0 mm, 54.0 mm, 40.5 mm SVL, respectively, Hiyare near Galle, 068060N, 808200E, 60 m, 9 March 2003, collected by S. Batuwita; WHT 7007, male, 73.0 mm SVL, Kanneliya Forest Reserve, 068150N, 808200E, 150 m, 13 July 1999, collected by S. Batuwita and I. Alagiyawadu; WHT 2370, female, 70.0 mm SVL, Morningside forest reserve, Sinharaja World Heritage Site, 06825010"N, 80836030"E, 500 m, 12 January 1999, collected by M. M. Bahir and S. Nanayakkara; WHT 6768, male, 34.0 mm SVL, same location data of WHT 2370, 14 January 1999; WHT 6959, 6977, two males, 48.0, 53.0 mm SVL, respectively, Udugama, Galle, 068140N, 808200E, 30 m, 10 February 2003, collected by P. Wickramatilake; NMSL CCA 1771, female, 71.2 mm SVL, no collection data [coordinates not correctly copyable from original paper] 
DiagnosisDiagnosis. Maximum SVL 73.0 mm; supranasals widely separated; postnasals absent; prefrontals widely separated, extended to lateral surface of snout; first supraocular not in contact with frontal; second supraocular in contact with frontal; frontoparietals paired, longer than wide; interparietal shorter than frontoparietal; parietal eye present; nuchals one pair; supraciliaries five, third distinctly elongate; pretemporals two, only upper contact with parietal; primary temporals two; secondary temporals two, separated by tertiary temporal scale, equal in size; supralabials seven; postsupralabials two; lower eyelid scaly; infralabials seven; first pair of chin shields in median contact; second pair of chin shields separated by a single median scale; dorsal body scale carination extends onto nuchals, parietals, supraoculars, and to temporals; paravertebrals 31–34; ventrals 46–52; longitudinal scale rows at midbody 28–32; keels on dorsal and lateral body scales 3–5; subdigital lamellae under fourth digit of pes 16–18; dorsolateral stripe narrow, a half-scale wide; dorsal coloration bronze brown with series of uninterrupt- ed dark brown longitudinal lines; and skin fragile.

Comparisons. “Here I compare E. greeri to congeners from Asia, listing only opposing suites of character states. Eutropis andama- nensis: six supraciliaries, primary temporal single; E. austini: first supraocular contact with frontal; interparietal as long as frontoparietal; 5–7 keeled dorsal body scales up to nuchals; dorsolateral stripe one and a half scale wide; skin not fragile, dorsum copper brown color without uninterrupted dark longitudinal lines; E. beddomei: prefrontals in contact medially, six supraciliaries; E. carinata: supraciliaries 6–7, maximum SVL 122.0 mm, first pair of chin shields separated by a single median scale; E. clivicola: prefrontals in contact medially, first pair of chin shields separated by a single median scale; E. dissimilis: supra- nasals in contact medially, prefrontals in contact medially, lower eyelid with a transparent window; E. bibronii: lower eyelid with a transparent window, subdigital lamellae under fourth digit of pes 20–22; E. floweri: first pair of chin shields separated by a single median scale, paravertebral scales 40–43; E. gansi: first pair of chin shields separated by a single median scale, both pretempo- rals in contact with parietal; E. innotata: lower eyelid with a transparent window, six supraciliaries; E. longicaudata: supra- nasals in contact medially, postnasal present; E. macularia: first pair of chin shields separated by a single median scale, both pretemporals in contact with parietal; E. madaraszi: single postsupralabial, well-developed median keels in all digits, copper brown dorsum coloration; E. multifasciata: prefrontals in contact medially, postnasal present; E. nagarjuni: two pairs of nuchals, prefrontals in contact medially; E. novemcarinata: supranasals in contact medially, lower eyelid with a transparent window; E. quadricarinata: prefrontals in contact medially, postnasal present; E. rugifera: 22–27 subdigital lamellae beneath fourth digit of pes; E. tammanna: prefrontal in contact medially, 37–40 paravertebrals, reddish gular and throat coloration, no dorsolateral stripe; E. trivittata: supranasals in contact medially, prefrontals in contact medially; E. tytleri: prefrontals in contact medially, postnasal present.” (from Batuwita 2016: 491)
 
CommentIllustrations: Eutropis greeri is illustrated by Somaweera and Somaweera (2009) in figs. 255A, B (identified as E. floweri) and fig. 257A (identified as E. madaraszi).

Sympatry: L. fallax, Lankascincus gansi, Lankascincus dorsicatenatus, Lankascincus greeri, E. carinata, and O. wiegmanni.

Habitat: rainforest areas (forest reserves) of the wet zone; terrestrial species, often observed in leaf litter in lowland rain forests. 
EtymologyThe species name is a patronym in the Latin genitive singular, in honor of Allen E. Greer, for his special commitment to the scincid fauna of the world and for providing evidence for undescribed Eutropis species from Sri Lanka (see Greer and Nussbaum, 2000). 
References
  • Batuwita, Sudesh 2015. Description of Two New Species of Eutropis (Reptilia: Scincidae) from Sri Lanka with a Redescription of Eutropis madaraszi (Méhely) Journal of Herpetology Sep 2016, Vol. 50, No. 3: 486-496. - get paper here
  • Das, Indraneil; de Silva, Anslem; Austin, Christopher C. 2008. A new species of Eutropis (Squamata: Scincidae) from Sri Lanka. Zootaxa 1700: 35-52 - get paper here
  • Somaweera, R. & Somaweera, N. 2009. Lizards of Sri Lanka: a colour guide with field keys. Chimaira, Frankfurt, 304 pp.
 
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