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Eutropis rugifera (STOLICZKA, 1870)

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Higher TaxaScincidae, Mabuyinae (Mabuyini), Scincoidea, Sauria, Squamata (lizards)
Subspecies 
Common NamesE: Rough-scaled Sun Skink
G: Nicobaren-Mabuye, Grobschuppiger Sonnenskink 
SynonymTiliqua rugifera STOLICZKA 1870: 107 or 170
Euprepes (Tiliqua) percarinatus PETERS 1871: 571
Euprepes (Tiliqua) percarinatus var. borneensis PETERS 1871: 572
Tiliqua percarinata var. borneensis — FISCHER 1885: 3
Mabuia rugifera — BOULENGER 1887: 184
Mabuia quinquecarinata WERNER 1896: 12 (fide SMITH 1935, TAYLOR 1963)
Mabuia rubricollis BARTLETT 1895: 87 (fide DE ROOIJ 1915: 160)
Mabuia quinquecarinata — WERNER 1910: 43
Mabuia quinquecarinata — DE ROOIJ 1915: 159
Mabuya rugifera — DE ROOIJ 1915: 160
Mabuya rugifera — DAMMERMAN 1929
Mabuya rugifera — SMITH 1935: 273
Mabuya rugifera — TAYLOR 1963: 947
Dasamia rugifera — HORTON 1973
Dasamia rugifera — DAS 1996: 45
Mabuya rugifera — MANTHEY & GROSSMANN 1997: 273
Mabuya rugifera — COX et al. 1998: 111
Mabuya rugifera — GREER & NUSSBAUM 2000
Eutropis rugifera — MAUSFELD et al. 2002
Mabuya rugifera — MALKMUS et al. 2002
Mabuya rugifera — DAS 2004
Eutropis rugifera — GRISMER 2011
Eutropis rugifera (?) — ONN et al. 2010
Eutropis rugifera — GRISMER & QUAH 2019 
DistributionIndia, S Thailand, Yala, Satun,
West Malaysia (Pulau Pangkor, Perak, Pahang, Selangor), Singapore,
Indonesia (Borneo, Sumatra, Pulau Nias, Mentawai Islands, Java, Bali, Natuna islands)
Philippine islands (Mindanao: Zamboanga Peninsula)

Type locality: Camorta, Nicobars

Euprepes percarinatus: type locality: Malang, East Java
Mabuia rubricollis: type locality: Kuching, Borneo
Mabuia quinquecarinata: type locality: Sumatra  
Reproductionviviparous (not imputed, fide Zimin et al. 2022) 
TypesHolotype: ZSI 2350 (apparently the single type mentioned by STOLICZKA 1870); the BMNH online catalogue lists BMNH 1946.8.18.22-24 as “types” without further details.
Syntypes: MSNG 28107 [borneensis] 
Diagnosis 
CommentDistribution: see map in Karin et al. 2017 (Fig. 1). The population from Mindanao is genetically divergent and may represent a different species ( Karin et al. 2017). Not listed as such by DAS 1996 (Biogeogr. SE-Asia). Has been reported from the Nicobar Islands but not listed by DAS 1999.

Animals from Bali lack the typical 5-7 longitudinal lines and are uniform brown dorsally and laterally. 
References
  • Amarasinghe, A.A. THASUN; NIKOLAY A. POYARKOV JR., PATRICK D. CAMPBELL, SANDY LEO, JATNA SUPRIATNA & JAKOB HALLERMANN 2017. Systematics of Eutropis rugifera (Stoliczka, 1870) (Squamata: Scincidae) including the redescription of the holotype. Zootaxa 4272 (1): 103–118 - get paper here
  • Baker, Nick 2016. Rough-scaled Sun Skink Eutropis rugifera (plain morph) at Panti Forest, Peninsular Malaysia. SEAVR 2016: 146-147 - get paper here
  • Boulenger, G. A. 1887. Catalogue of the lizards in the British Museum (Nat. Hist.) III. Lacertidae, Gerrhosauridae, Scincidae, Anelytropsidae, Dibamidae, Chamaeleontidae. London: 575 pp. - get paper here
  • Boulenger, George A. 1890. The Fauna of British India, Including Ceylon and Burma. Reptilia and Batrachia. Taylor & Francis, London, xviii, 541 pp. - get paper here
  • Capocaccia, L. 1961. Catalogo dei tipi di Rettili del Museo Civico di Storia Naturale di Genova [MSNG]. Ann. Mus. Civ. Stor. Nat. Giacomo Doria 72: 86-111 - get paper here
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Cox, Merel J.; Van Dijk, Peter Paul; Jarujin Nabhitabhata & Thirakhupt,Kumthorn 1998. A Photographic Guide to Snakes and Other Reptiles of Peninsular Malaysia, Singapore and Thailand. Ralph Curtis Publishing, 144 pp.
  • Dammerman, K.W. 1929. On the zoogeography of Java. Treubia 11: 1-88 - get paper here
  • Das , I. 2004. Lizards of Borneo. Natural History Publications, Kota Kinabalu, Borneo
  • Das, I. 1999. Biogeography of the amphibians and reptiles of the Andaman and Nicobar Islands, India. In: Ota,H. (ed) Tropical Island herpetofauna..., Elsevier, pp. 43-77
  • Das, INDRANEIL, BASUDEB DATTAGUPTA and NEMAI CHARAN GAYEN. 1998. History and catalogue of reptile types in the collection of the Zoological Survey of India. J. south Asian nat. Hist. 3 (2): 121-172
  • Davis, Hayden R.; L. Lee Grimser, Anthony J. Cobos, Matthew L. Murdoch, Alexandra X. Sumarli, Sharual Anuar, M. A. Muin, Perry Wood, Jr., Evan S. H. Quah 2018. Checklist of the Herpetofauna of Hutan Lipur Gunung Senyum, Pahang, Peninsular Malaysia. Russ. J. Herpetol. 25 (3): 207-220 - get paper here
  • de Rooij, N. de 1915. The Reptiles of the Indo-Australian Archipelago. I. Lacertilia, Chelonia, Emydosauria. Leiden (E. J. Brill), xiv + 384 pp. - get paper here
  • FIGUEROA, ALEX; MARTYN E.Y. LOW, KELVIN K.P. LIM 2023. Singapore’s herpetofauna: updated and annotated checklist, history, conservation, and distribution. Zootaxa 5287 (1): 001–378 - get paper here
  • Fischer, J.G. 1886. Ueber eine Kollektion Reptilien und Amphibien von der Insel Nias und über eine zweite Art der Gattung Anniella GRAY. Abh. Naturw. Ver. Hamburg 19 (1): 3-10
  • Greer,A.E. & NUSSBAUM,R.A. 2000. New character useful in the systematics of the scincid lizard genus Mabuya. Copeia 2000 (2): 615-618 - get paper here
  • GRISMER, L. LEE; & EVAN S. H. QUAH 2019. An updated and annotated checklist of the lizards of Peninsular Malaysia, Singapore, and their adjacent archipelagos. Zootaxa 4545 (2): 230–248 - get paper here
  • Grismer, L.L. 2011. Lizards of Peninsular Malaysia, Singapore and their adjacent archipelagos. Edition Chimaira, Frankfurt, 728 pp. [review in Herp. Rev. 43: 155] - get paper here
  • Herlambang, A. E., Riyanto, A., Munir, M., Hamidy, A., Kimura, K., Eto, K., & Mumpuni, M. 2022. AFTER 16 YEARS: AN UPDATE CHECKLIST OF HERPETOFAUNA ON THE NATUNA ISLANDS, INDONESIA. TREUBIA, 49(2), 67-84 - get paper here
  • Horton, David Robert 1973. A New Scincid Genus from Southeast Asia. [Dasamia]. Journal of Herpetology 7 (3): 283-298 - get paper here
  • Karin BR, Das I, Jackman TR, Bauer AM. 2017. Ancient divergence time estimates in Eutropis rugifera support the existence of Pleistocene barriers on the exposed Sunda Shelf. PeerJ 5:e3762 - get paper here
  • Kurniawan, N., Septiadi, L., Fathoni, M., & Muammar, A. 2022. A Checklist of the Herpetofauna of Nusa Kambangan Island, Central Java, Indonesia. Tropical Life Sciences Research, 33(2): 91–131 - get paper here
  • Malkmus, R.; Manthey, U.; Vogel, G. Hoffmann, P. & Kosuch, J. 2002. Amphibians and reptiles of Mount Kinabalu (North Borneo). A.R.G. Ganther Verlag, Rugell, 404 pp.
  • Manthey, U. & Grossmann, W. 1997. Amphibien & Reptilien Südostasiens. Natur und Tier Verlag (Münster), 512 pp. - get paper here
  • MAUSFELD, Patrick & Andreas SCHMITZ 2003. Molecular phylogeography, intraspecific variation and speciation of the Asian scincid lizard genus Eutropis Fitzinger, 1843 (Squamata: Reptilia: Scincidae): taxonomic and biogeographic implications. Org. Divers. Evol. 3: 161–171 - get paper here
  • Milto, Konstantin D.; Yury A. Lukin 2020. A Revised Herpetofauna of Ujung Kulon National Park, West Java, Indonesia. Russian Journal of Herpetology 27 (6): 353-368 - get paper here
  • Onn, Chan Kin; J. van Rooijen, L. Lee Grismer, Daicus Belabut,, Mohd. Abdul Muin Md. Akil, Hamidi Jamaludin, Rick Gregory, and Norhayati Ahmad 2010. FIRST REPORT ON THE HERPETOFAUNA OF PULAU PANGKOR, PERAK, MALAYSIA. Russ. J. Herpetol. 17 (2): 139 – 146 - get paper here
  • Peters, W. 1871. Uber neue Reptilien aus Ostafrika und Sarawak (Borneo), vorzüglich aus der Sammlung des Hrn. Marquis J. Doria zu Genua. Mber. k. preuss. Akad. Wiss., Berlin [1871]: 566-581 - get paper here
  • Smith, M.A. 1926. “Spolia Mentawia” - Reptiles and Amphibians. Ann. Mag. Nat. Hist. (9) 18: 76-81 - get paper here
  • Smith, M.A. 1935. The fauna of British India, including Ceylon and Burma. Reptiles and Amphibia, Vol. II. Sauria. Taylor and Francis, London, 440 pp.
  • Smith,M.A. 1935. On a collection of reptiles and amphibians from Perak, Malay Peninsula. Bull. Raffles Mus. No 10: 61-63
  • Stoliczka, F. 1870. Observations of some Indian and Malayan Amphibia and Reptilia (abstract). Ann. Mag. Nat. Hist. (4) 6: 105-109 - get paper here
  • Stoliczka, F. 1870. Observations on some Indian and Malayan Amphibia and Reptilia. J. Asiat. Soc. Bengal 39: 134-228. - get paper here
  • Sworder, G.H. 1933. Notes on some reptiles from the Malay Peninsula. Bull. Raffles Mus. No 8: 101-105
  • Taylor, E.H. 1963. The lizards of Thailand. Univ. Kansas Sci. Bull. 44: 687-1077. - get paper here
  • Teo, R.C.H. & N.J. Thomas 2019. Updated Inventory of Amphibians, Reptiles and Mammals of the Bukit Timah Nature Reserve, Singapore. Gardens’ Bulletin Singapore 71(Suppl. 1):145-183; doi: 10.26492/gbs71(suppl.1).2019-06 - get paper here
  • Teo, R.C.H. & Rajathurai, S. 1997. Mammals, reptiles and amphibians in the Nature Reserves of Singapore - diversity, abundance and distribution. Proc. Nature Reserves Survey Seminar. Gardens’ Bulletin Singapore 49: 353-425
  • TEYNIÉ, ALEXANDRE; PATRICK DAVID, & ANNEMARIE OHLER 2010. Note on a collection of Amphibians and Reptiles from Western Sumatra (Indonesia), with the description of a new species of the genus Bufo. Zootaxa 2416: 1–43 - get paper here
  • Venugopal, P.D. 2010. An updated and annotated list of Indian lizards (Reptilia: Sauria) based on a review of distribution records and checklists of Indian reptiles. Journal of Threatened Taxa 2 (3): 725-738. - get paper here
  • Werner, F. 1896. Zweiter Beitrag zur Herpetologie der indo-orientalischen Region. Verh. Zool. Bot. Ges. Wien 46: 6-24 - get paper here
  • Werner,F. 1910. Über neue oder seltene Reptilien des Naturhistorischen Museums in Hamburg. ii. Eidechsen. Jahrb. Hamburg. Wiss. Anst., vol. 27 (1909), suppl. no. 2, 1910, pp. 1-46; reprinted: 1910, Mitteil. Naturhist. Mus. Hamburg, vol. 27: 205-) - get paper here
  • Zimin, A., Zimin, S. V., Shine, R., Avila, L., Bauer, A., Böhm, M., Brown, R., Barki, G., de Oliveira Caetano, G. H., Castro Herrera, F., Chapple, D. G., Chirio, L., Colli, G. R., Doan, T. M., Glaw, F., Grismer, L. L., Itescu, Y., Kraus, F., LeBreton 2022. A global analysis of viviparity in squamates highlights its prevalence in cold climates. Global Ecology and Biogeography, 00, 1–16 - get paper here
 
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