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Gehyra granulum DOUGHTY, PALMER, BOURKE, TEDESCHI, OLIVER & MORITZ, 2018

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common NamesKimberley granular-toed gecko 
SynonymGehyra granulum DOUGHTY, PALMER, BOURKE, TEDESCHI, OLIVER & MORITZ in DOUGHTY et al. 2018: 220 
DistributionAustralia (Western Australia)

Type locality: Bell Gorge water hole, Western Australia (16.99333°S; 125.20489°E).  
Reproduction 
TypesHolotype: WAM R176212 (CCM1270) (male), collected on 22 May 2013 by G. Bourke, J. Bragg, R. Laver and P. Title. Paratypes (10). WAM R158823 (male), Bigge Island (14.6019°S, 125.2039°E); WAM R168558 (BP01286) (female), Champagny Island (15.30°S, 124.25°E); WAM R168561 (BP01272) (female), Byam Martin Island (15.3833°S, 124.35°E); WAM R171072 (BP00315) (male), Augustus Island (15.3900°S, 124.5931°E); WAM R171403 (male), Bachsten Creek bush camp (15.9897°S, 125.3294°E); WAM R171426 (female), Prince Regent River National Park (15.9810°S, 125.3731°E); WAM R176224 (CCM1339) (male), Gibb River Road, roadside (17.1189°S, 125.1329°E); WAM R176245 (CCM1492) (male), Gibb River Road (17.16653°S, 125.34761°E); NMV D75778 (male) and NMV D75774 (female), western end of Artesian Range (16.54834°S, 124.9961°E). 
DiagnosisDiagnosis. A Gehyra with small body size (average 40 mm, 35–46 mm SVL), no flap of skin between limbs, dorsal half of rostral deeply furrowed with groove, 1 internasal scale present or absent, 2 postnasals of similar size, first supralabial slightly taller and narrower than second, 2 pairs of chin shields, snout short and convex in lateral view, first digit of manus and pes without claw, 5 or 6 divided subdigital lamellae on fourth toe, proximal lamellae usually separated by 1–3 granular scales (10% lack granules), mode of 15 (range 10–19) pre-cloacal pores in males arranged in shallow chevron pointing anteriorly. Background colouration in life dull tan to brown with scattered small pale spots and larger dark blotches or irregular lines on dorsum, often forming short transverse rows.

Comparisons with other species. Gehyra granulum sp. nov. is most similar to other small-bodied Gehyra from the Kimberley. It differs from G. nana, G. girloorloo and G. pluroporosa sp. nov. by usually possessing 1–3 small granules on the proximal subdigital lamellae and often lacks an internarial scale. From G. spheniscus, another small-bodied Gehyra with granules on the digits, it differs by number of granules on proximal lamellae (1–3 vs a small wedge of 5–7; Fig. 14), fewer pre-cloacal pores in males (10–19 vs 21–27) and different colouration (reddish-brown with large dark and small pale spots vs purplish-brown with similar-sized dark and pale spots). 
CommentHabitat. This species has been collected from low rocks and rock faces in sandstone rock formations in open savannah woodlands with spinifex. Occasionally collected from trees, but only when in association with rocky outcrops. It is broadly sympatric with the larger-bodied species G. multiporosa, G. occidentalis, and G. pseudopunctata sp. nov. and with the more similarly sized G. spheniscus in the southwest Kimberley, and overlaps geographically with G. kimberleyi and true G. nana in the southern and eastern Kimberley.

Distribution: see map in Doughty et al. 2018: 203 (Fig. 1). 
Etymologygranulum (Latin) refers to the granular scales between the proximal subdigital lamellae that characterises this species. 
References
  • DOUGHTY, PAUL; GAYLEEN BOURKE, LEONARDO G. TEDESCHI, RENAE C. PRATT, PAUL M. OLIVER, RUSSELL A. PALMER, CRAIG MORITZ 2018. Species delimitation in the Gehyra nana (Squamata: Gekkonidae) complex: cryptic and divergent morphological evolution in the Australian Monsoonal Tropics, with the description of four new species Zootaxa 4403 (2): 201–244 - get paper here
  • Ellis, Ryan J.; Paul Doughty and Aaron M. Bauer 2018. An annotated type catalogue of the geckos and pygopods (Squamata: Gekkota: Carphodactylidae, Diplodactylidae, Gekkonidae, Pygopodidae) in the collection of the Western Australian Museum Records of the Western Australian Museum 33: 051–094 - get paper here
 
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