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Gloydius intermedius (STRAUCH, 1868)

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Higher TaxaViperidae, Crotalinae, Colubroidea, Alethinophidia, Serpentes, Squamata (snakes) 
Common NamesE: Central Asian pitviper 
SynonymTrigonocephalus intermedius STRAUCH 1868
Agkistrodon halys intermedius STRAUCH 1876
Halys intermedia — PETERS 1877: 735
Ancistrodon intermedius — BOULENGER 1890: 140
Ancistrodon intermedius — BOULENGER 1896: 525
Agkistrodon blomhoffi intermedius — STEJNEGER 1907: 464
Agkistrodon halys cognatus GLOYD 1977
Agkistrodon halys intermedius — HARDING & WELCH 1980
Agkistrodon intermedius intermedius — WELCH 1994: 11
Ancistrodon blomhoffi intermedius — DESPAX 1913: 181
Agkistrodon intermedius — ORLOV & BARABANOV 1999
Gloydius intermedius — MCDIARMID, CAMPBELL & TOURÉ 1999: 306
Gloydius intermedius — GUMPRECHT et al. 2004
Gloydius intermedius — WALLACH et al. 2014: 311 
DistributionN China (incl. Liaoning, Jilin, Heilongjiang),
E Russia, North Korea, South Korea

Type locality: Irkutsk, Eastern Siberia; Terra typica restricta (hoc loco ). Malyi Hinggan Mountain Range (Xiao Hinggan Ling), Cape Tyr on the Amur River and Cape Kofel on the Ussuri River, Russian Far East Region. Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
TypesLectotype. ZISP 2221. Prom. Tyr (Amur). Leg.: Schrenck, 1855 

Synonymy partly after KHALIKOV & ANANJEVA (pers. comm.).

Distribution: see map in ORLOV & BARABANOV 1999.

Diagnosis. Total length up to 800 mm and more. Dorsal scales in 23 (rarely 21) rows at midbody. Supralabials 7 – 8; ventrals 148 – 175; subcaudals 34–52.

Diagnosis. A large viper of the genus, up to at least 800 mm total length. Supralabial scales 7-8. Dorsal scales in 23, very rarely 21, rows around midbody; ventral scales between 148- 175; subcaudal scales paired, between 34-52. Body with 28-45 dark transverse bands, each 3- 6 scales wide and extending down to scale row 2 or 1 [WAGNER et al. 2015]. 
  • Adnagulov E.V. 2016. On the distribution of amphibians and reptiles in the Evreiskaya Autonomous Oblast' (Russia). [in Russian] Current Studies in Herpetology, 16(3/4): 87–106 - get paper here
  • Boulenger, G.A. 1896. Catalogue of the snakes in the British Museum, Vol. 3. London (Taylor & Francis), xiv + 727 pp. - get paper here
  • Boulenger, George A. 1890. A list of the reptiles and batrachians of Amoorland [Russia]. Ann. Mag. nat. Hist. (6) 5: 137-144 - get paper here
  • Chen, Y.; Zhang, D.; Jiang, K. & Wang, Z. 1992. Evaluation of Snake Venoms among Agkistrodon Species in China. Asiatic Herpetological Research 4: 58-61 - get paper here
  • Despax, RAYMOND 1913. Sur une collection de reptiles et de batraciens rassemblée par M. le Dr. Legendre dans les marches Thibétaines. Bull. Mus. Hist. Nat., Paris, [ser. 1], 19 (4): 179-183 - get paper here
  • Gumprecht, A.; Tillack, F.; Orlov, N.L.; Captain, A. & Ryabow, S. 2004. Asian pitvipers. Geitje Books, Berlin, 368 pp.
  • Guo, P.; Zhang, F.J. & Chen, Y.Y. 1999. The Hemipenes of Chinese Species of Deinagkistrodon and Gloydius (Serpentes: Crotaiinae). Asiatic Herpetological Research 8: 38-42 - get paper here
  • Kharin V E 2011. Annotated catalogue of amphibians and reptiles (Amphibia, Reptilia) of the Far-Eastern Marine Biosphere FEB RAS [in Russian]. Biodiversity and Environment of Far East Reserves, Vladivostok, DVMBGPZ DVO RAN 2011 (1): 30-48
  • LIN Xuan-Long, GUO Dong-Sheng 2012. A New Record of Snake in Beijing: Gloydius intermedius. Chinese Journal of Zoology 2012, 47(2):136-137 - get paper here
  • Macey, J. Robert;Papenfuss, Theodore J.;Zhao, Ermi 1988. The snakes of Ningxia Hui autonomous region as an indication of a herpetofaunal corridor. Chinese Herpetological Research 2 (1): 4-5
  • McDiarmid, R.W.; Campbell, J.A. & Touré,T.A. 1999. Snake species of the world. Vol. 1. Herpetologists’ League, 511 pp.
  • Orlov, Nikolai L. and Andrei V. Barabanov. 1999. Analysis of nomenclature, classification, and distribution of the Agkistrodon halys - Agkistrodon intermedius complexes: a critical review. Russ. J. Herpetol. 6 (3): 167-192 - get paper here
  • Peters, W. 1877. Eine Übersicht über die während der sibirischen Expedition von 1876 von Hrn. Dr. O. Finsch gesammelten Säugethiere, Amphibien und Fische. Monatsberichte der Königlich Preussischen Akademie der Wissenschaften zu Berlin 1877: 734-738 - get paper here
  • Stejneger, LEONHARD H. 1907. Herpetology of Japan and adjacent territory. Bull. US Natl. Mus. 58: xx, 1-577 - get paper here
  • Strauch, A. 1868. Concerning poisonous snakes distributed in Russia. [in Russian]. Trudy Perv. Siezda Russ. Yestestv. Zool., 1: 294 (also cited as Trans. First Conf. Russ. Nat., St.Petersburg, 1: 271-297)
  • Toriba, Michihisa. 1999. Comments on the taxonomy of Agkistrodon listed in Stejneger (1907). Bulletin of the Herpetological Society of Japan 1999 (1): 3-5. - get paper here
  • Wagner, Philipp; Tiutenko, Arthur; Mazepa, Glib; Borkin, Leo J.; Simonov, Evgeniy 2015. Alai! Alai! – a new species of the Gloydius halys (Pallas, 1776) complex (Viperidae, Crotalinae), including a brief review of the complex. Amphibia-Reptilia 37 (published 31 Dec 2015) DOI: 10.1163/15685381-00003026 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
  • XIONG Ye, DING Li 2012. A Taxonomic Status of Fu Snakes in Mt. Wuling, Xinglong County, Hebei Province. Sichuan Journal of Zoology 31 (5) - get paper here
  • XU, Yan; Qin LIU, Edward A MYERS, , Lian WANG, Song HUANG, Yun HE,Peihao PENG and Peng GUO<br /> 2012. Molecular Phylogeny of the Genus Gloydius (Serpentes: Crotalinae). Asian Herpetological Research 3 (2): 127–132 - get paper here
  • ZHAO Wenge, LIU Xiaolong, & GUO Yumin 2000. Discovery of Gloydius intermedius in Heilongjiang Province. Sichuan Journal of Zoology (2000) 19 (3): 157-158 - get paper here
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • Zhao,E. & Adler,K. 1993. Herpetology of China. SSAR, Oxford/Ohio, 1-522
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