Goggia gemmula (BAUER, BRANCH & GOOD, 1996)

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Higher Taxa	Gekkonidae, Gekkota, Sauria, Squamata (lizards: geckos)
Subspecies
Common Names	E: Richtersveld Dwarf Leaf-toed Gecko
Synonym	Phyllodactylus gemmulus BAUER, BRANCH & GOOD 1996 Goggia gemmula — BAUER et al. 1997 Goggia gemmula — RÖSLER 2000: 83 Goggia gemmula — BATES et al. 2014: 109
Distribution	Republic of South Africa (Richtersveld), S Namibia (Karas District) Type locality: 22.6 km by road E Sendelingsdrif (Park Headquarter), Richtersveld National Park, Northern Cape Province, South Africa.
Reproduction	oviparous
Types	Holotype: CAS 193645, paratypes: PEM
Diagnosis	Diagnosis: A small species of the P. lineatus group, having an adult snout-vent length (SVL) of 23-30 mm and distinguished from other species in the genus and species group by the following combination of characteristics: head depressed; dorsal body scalation homogeneous; midbody scale rows 70-86; 4 preanal pores in males; dorsal color pattern with transverse rows of large yellow spots on a brown or mottled background. Phyllodactylus gemmulus differs from P . microlepidotus, P . porphyreus, and an undescribed species from the Swartberg Mountains by its smaller size, and from the last species and P. peringueyi by its homogeneous body scalation. It is distinguished from P. lineatus and several related forms (resurrected or described by Branch et aI., 1995) by its elongate trunk and its color pattern of large, diffuse yellow spots (small pale spots may be found in other species and small orange spots are seen in P. rupicolus from Namaqualand). It is distinguished from P. hexaporus and from P. lineatus by its lower number of preanal pores, and from all members of the group except P.linealus by a relatively low number of midbody scale rows. Phyllodaclylus gemmulus is also distinguished by four unique allozyme alleles (Bauer et al. 1996: 4)
Comment
Etymology	Named after Latin gemmula = a small jewel or gem, referring to the small size and bright (for the genus) dorsal coloration of the species.
References	Anonymous 1997. At home in the rocks, a new gecko emerges. National Geographic 191(6):unnumbered Bates, M.F.; Branch, W.R., Bauer, A.M.; Burger, M., Marais, J.; Alexander, G.J. & de Villliers, M.S. (eds.) 2014. Atlas and Red List of the Reptiles of South Africa, Lesotho, and Swaziland. Suricata 1. South African National Biodiversity Institute, Pretoria, 512 pp. Bauer A M. Good D A. Branch W R. 1997. The taxonomy of the Southern African leaf-toed geckos (Squamata: Gekkonidae), with a review of Old World "Phyllodactylus" and the description of five new genera. Proc. Cal. Acad. Sci. 49 (14): 447-497. - get paper here Bauer, A.M., and Branch, W.R. 2003. The herpetofauna of the Richtersveld National Park, Northern Cape Province, Republic of South Africa. Herpetological Natural History 8: 111-160 [2001] Bauer, A.M., W.R. Branch & D.A. Good 1996. A new species of rock-dwelling Phyllodactylus (Squamata: Gekkonidae) from the Richtersveld, South Africa. Occ. Pap. Mus. Nat. Sci. Louisiana State Univ. 71: 2-14. - get paper here Bauer, Aaron 1999. An Interview with Aaron Bauer. Gekko 1 (1): 16-21 CONRADIE, WERNER; WILLIAM R. BRANCH, & GILLIAN WATSON 2019. Type specimens in the Port Elizabeth Museum, South Africa, including the historically important Albany Museum collection. Part 2: Reptiles (Squamata). Zootaxa 4576 (1): 001–045 - get paper here Herrmann, H.-W.; W.R. Branch 2013. Fifty years of herpetological research in the Namib Desert and Namibia with an updated and annotated species checklist. Journal of Arid Environments 93: 94–115 - get paper here Rösler, H. 2000. Kommentierte Liste der rezent, subrezent und fossil bekannten Geckotaxa (Reptilia: Gekkonomorpha). Gekkota 2: 28-153
External links	IUCN Red List National Center for Biotechnology Information Google images

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