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Grayia tholloni MOCQUARD, 1897

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Higher TaxaColubridae, Grayiinae, Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
Subspecies 
Common NamesTholloni's African Water Snake 
SynonymGrayia tholloni MOCQUARD 1897: 11
Grayia fasciata BOULENGER 1901: 2 (fide LOVERIDGE 1957: 268)
Grayia tholloni — SCHMIDT 1923: 95
Grayia tholloni — LOVERIDGE 1936: 33
Grayia tholloni — BROADLEY 1998
Grayia tholloni — CHIRIO & INEICH 2006
Grayia tholloni — TRAPE & BALDÉ 2014
Grayia tholloni — WALLACH et al. 2014: 316 
DistributionS Sudan, Uganda, Kenya, Tanzania, N/E/S Democratic Republic of the Congo (Zaire), Congo, Angola, Central African Republic, Ethiopia, Gambia, Senegal (HR 26: 157; HR 27: 88), Nigeria, Guinea (Conakry), possibly Cameroon (Perret 1961), Zambia, S Chad

Type locality: Congo (Brazzaville). Map legend:
TDWG region - Region according to the TDWG standard, not a precise distribution map.

NOTE: TDWG regions are generated automatically from the text in the distribution field and not in every cases it works well. We are working on it.
 
Reproductionoviparous 
TypesSyntypes: MT (2) and MNHN (2) (Thollon), largest specimen is 810 mm. 
CommentAn almost entirely aquatic species.

Not in Gabon fide PAUWELS & VANDE WEGHE 2008. 
Etymologynamed after François-Romain Thollon, a botanist at the Muséum d’histoire naturelle (Paris) and the collector of the types. 
References
  • Barnett, Linda K. 2001. The herpetofauna of Abuko Nature Reserve, the Gambia. Herpetological Bulletin (77): 5-1 - get paper here
  • Barnett, Linda K. & Emms, Craig 2005. Common reptiles of The Gambia. Rare Repro, Hailsham, East Sussex, 24 pp.
  • Böhme, Wolfgang, Mark-Oliver Rödel, Christian Brede & Philipp Wagner 2011. The reptiles (Testudines, Squamata, Crocodylia) of the forested southeast of the Republic Guinea (Guinée forestière), with a country-wide checklist. Bonn zoological Bulletin 60 (1): 35-61 - get paper here
  • Boulenger, G. A. 1901. Materiaux pour la faune du Congo. Batraciens et reptiles nouveaux. Ann. Mus. Congo Belge, Zool. (sect. C. ser. 1) 2: 7-14.
  • Broadley, D. G. & HOWELL, K. M. 1991. A check list of the reptiles of Tanzania, with synoptic keys. Syntarsus 1: 1—70
  • Broadley, D.G. 1983. Records of two species of Grayia (Serpentes: Colubridae) from Zambia. Black Lechwe (n.s.) 5: 21-25
  • Broadley, D.G. 1991. The Herpetofauna of Northern Mwinilunga Distr., Northw. Zambia. Arnoldia Zimbabwe 9 (37): 519-538
  • Broadley, D.G. 1998. The reptilian fauna of the Democratic Republic of the Congo (Congo-Kinshasa). In: Schmidt, K.P. and Noble, G.K., Contributions to the Herpetology of the Belgian Congo... [reprint of the 1919 and 1923 papers]. SSAR Facsimile reprints in Herpetology, 780 pp.
  • Broadley, D.G.; Doria, C.T. & Wigge, J. 2003. Snakes of Zambia. An Atlas and Field Guide. Edition Chimaira, Frankfurt, 280 pp. [review in Sauria 26 (3): 21]
  • Chifundera, K. 1990. Snakes of Zaire and their bites. Afr. Stud. Monogr. (Kyoto) 10(3): 137-157.
  • Chirio, L. & Lebreton, M. 2007. Atlas des reptiles du Cameroun. MNHN, IRD, Paris 688 pp.
  • Chirio, Laurent and Ivan Ineich 2006. Biogeography of the reptiles of the Central African Republic. African Journal of Herpetology 55(1):23-59. - get paper here
  • Haagner,G.V.; Branch,W.R. & Haagner,A.J.F. 2000. Notes on a collection of reptiles from Zambia and adjacent areas of the Democratic Republic of the Congo. Annals of the Eastern Cape Museum 1: 1 – 25
  • Hughes, B. 1983. African snake faunas. Bonner Zoologische Beiträge 34: 311-356 - get paper here
  • Largen, M.J.; Spawls, S. 2010. Amphibians and Reptiles of Ethiopia and Eritrea. Edition Chimaira, Frankfurt, 694 pp.
  • Largen,M.J. & Rasmussen,J.B. 1993. Catalogue of the snakes of Ethiopia (Reptilia Serpentes), including identification keys. Tropical Zoology 6: 313-434 - get paper here
  • Loveridge, A. 1936. African reptiles and amphibians in the Field Museum of Natural History. Zool. Ser. Field Mus. Nat. Hist., Chicago, 22 (1): 1-122 - get paper here
  • Loveridge, Arthur 1929. East African reptiles and amphibians in the United States National Museum. Bull. US Natl. Mus. (151): 1-135 - get paper here
  • Loveridge,A. 1957. Check list of the reptiles and amphibians of east Africa (Uganda, Kenya, Tanganyika, Zanzibar). Bull. Mus. Comp. Zool. Harvard 117 (2): 153-362 - get paper here
  • Mocquard, M.F. 1897. Sur une collection de Reptiles recueillis par M. Haug, à Lambaréné. Bull. Soc. Philom. Paris, (8) 9: 5-20 - get paper here
  • Pauwels,O. 1996. Geographic distribution: Grayia tholloni. Herpetological Review 27 (2): 88 - get paper here
  • Perret 1961. Bull. Soc. neuchâtel. Sci. nat., 84: 133.
  • Pitman,C.R.S. 1974. A guide to the snakes of Uganda. Codicote, Wheldon & Wesley, L., 290 pp.
  • Schmidt, K. P. 1923. Contributions to the herpetology of the Belgian Congo based on the collection of the American Museum Congo Expedition, 1909-1915. Part II. Snakes, with field notes by Herbert Lang and James P. Chapin. Bull. Amer. Mus. nat. Hist. 49 (1): 1-146
  • Spawls, S.; Howell, K.; Drewes, R.C. & Ashe, J. 2002. A field guide to the reptiles of East Africa. Academic Press, 543 pp. [reviews in HR 34: 396 and Afr. J. Herp. 51; 147] - get paper here
  • Trape, J.-F. & Mané, Y. 2006. Guide des serpents d’Afrique occidentale. Savane et désert. [Senegal, Gambia, Mauritania, Mali, Burkina Faso, Niger]. IRD Editions, Paris, 226 pp. - get paper here
  • Trape, J.F. & R. ROUX-ESTÈVE 1995. Les serpents du Congo: liste commentée et clé de détermination. Journal of African Zoology 109 (1): 31-50
  • TRAPE, JEAN-FRANÇOIS & CELLOU BALDÉ 2014. A checklist of the snake fauna of Guinea, with taxonomic changes in the genera Philothamnus and Dipsadoboa (Colubridae) and a comparison with the snake fauna of some other West African countries. Zootaxa 3900 (3): 301–338
  • Trape, Jean-François; Mané, Youssouph 1995. Grayia tholloni. Herpetological Review 26 (3): 156 - get paper here
  • Trape, Jean-François; Mané, Youssouph 2000. Les serpents des environs de Dielmo (Sine-Saloum, Sénégal). Bull. Soc. Herp. France 95: 19-35 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. Taylor and Francis, CRC Press, 1237 pp.
  • Witte, G. F. de 1933. Reptiles récoltés au Conge Belge par le Dr. H. Schouteden et par M. G.-F. Witte. Ann. Mus. Conge belge Zool. Ser. 1 Tome III: 53-100.
 
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