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Hebius atemporalis (BOURRET, 1934)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Tonkin Keelback
Chinese: 无颞鳞腹链蛇 
SynonymNatrix atemporalis BOURRET 1934: 3
Natrix atemporalis — BOURRET 1935: 2
Natrix atemporalis — SMITH 1943: 287
Amphiesma atemporalis — MALNATE 1960
Amphiesma atemporalis — ZHAO 2006
Amphiesma atemporale — DAVID et al. 2007
Hebius atemporale — GUO et al. 2014
Amphiesma atemporale — WALLACH et al. 2014: 28
Hebius atemporalis — ZHOU et al. 2019 
DistributionN Vietnam,
China (Hong Kong, Guangdong, and Guangxi west to Yunnan and Guizhou)

Type locality: Ton-king (= North Vietnam)  
Reproductionoviparous 
TypesSyntypes: MNHN-RA 1935.0077-0078, MNHN-RA 1935.0450, MNHN-RA 1938.0119-0120 (5 syntypes); perhaps MNHN-RA 1958.0461-0462 are also syntypes? 
Diagnosis 
Comment 
References
  • Bourret, R. 1934. Notes herpétologiques sur l'Indochine Française VI-VII. Bull. Gén. Instr. Publique, Hanoi, 1934 (4): 73-84
  • Bourret,R. 1935. Notes herpétologiques sur l'Indochine Française X. Les serpents de la station d’altitude du Tam-dao. Bull. Gen. Instr. Pub. Hanoi 8 (avril): 259-271 (1-13)
  • David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here
  • Francis, A. 2021. A field guide to the snakes of Hong Kong. Regal Printing, Hong Kong - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Malnate, E.V. & Romer, J.D. 1969. A snake new to the fauna of Hong Kong and China: Amphiesma atemporalis (Bourret). Notulae Naturae (Philadelphia) 424: 1–8.
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Smith, M.A. 1943. The Fauna of British India, Ceylon and Burma, Including the Whole of the Indo-Chinese Sub-Region. Reptilia and Amphibia. 3 (Serpentes). Taylor and Francis, London. 583 pp.
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56 - get paper here
 
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