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Hebius bitaeniatus (WALL, 1925)

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Higher TaxaColubridae (Natricinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes)
Subspecies 
Common NamesE: Kutkai Keelback
Chinese: 黑带腹链蛇 
SynonymNatrix bitaeniata WALL 1925: 806
Natrix bitaeniata — MELL 1931 [1929]
Amphiesma bitaeniata MALNATE 1960
Amphiesma bitaeniatum — DAVID, VOGEL & PAUWELS 1998
Amphiesma sp. — CHAN-ARD et al. 1999: 155 (fide DAVID & PAUWELS)
Amphiesma bitaeniatum — DAVID et al. 2007
Hebius bitaeniatum — GUO et al. 2014
Amphiesma bitaeniatum — WALLACH et al. 2014: 29
Amphiesma bitaeniatum — CHAN-ARD et al. 2015: 233
Hebius bitaeniatus — HIKIDA 2015
Hebius bitaeniatus — ZHOU et al. 2019 
DistributionBurma (Myanmar), Thailand, Vietnam (Yen Bai etc.),
China (W Yunnan, Hunan, Guizhou), Laos (Xiangkhoang)

Type locality: “Kutkai, North Shan States, Burma, 6000 feet”  
Reproductionoviparous 
TypesSyntypes: BMNH 1946.1.13.58, 
Diagnosis 
CommentDAVID & PAUWELS (2000) identified Amphiesma sp. — CHAN-ARD et al. 1999 in a book review in the Russ. J. Herpetol. 7: 89. 
EtymologyNamed after “bi-” = two or dual and Latin “taenia” = band or stripe. 
References
  • Chan-ard, T., Parr, J.W.K. & Nabhitabhata, J. 2015. A field guide to the reptiles of Thailand. Oxford University Press, NY, 352 pp. [see book reviews by Pauwels & Grismer 2015 and Hikida 2015 for corrections] - get paper here
  • Chan-ard,T.; Grossmann,W.; Gumprecht,A. & Schulz,K. D. 1999. Amphibians and reptiles of peninsular Malaysia and Thailand - an illustrated checklist [bilingual English and German]. Bushmaster Publications, Würselen, Gemany, 240 pp. [book review in Russ. J Herp. 7: 87] - get paper here
  • Das, I. 2012. A Naturalist's Guide to the Snakes of South-East Asia: Malaysia, Singapore, Thailand, Myanmar, Borneo, Sumatra, Java and Bali. Oxford J, ohn Beaufoy Publishing - get paper here
  • David P., G. Vogel & Pauwels,O. 1998. Amphiesma optatum (Hu & Djao, 1966) (Serpentes, Colubridae): an addition to the snake fauna of Vietnam, with a list of the species of the genus Amphiesma and a note on its type species. Journal of Taiwan Museum 51 (2): 83-92 - get paper here
  • David, P., Bain, R. H., Nguyen Quang Truong, Orlov, N. L., Vogel, G., Vu Ngoc Thanh & T. Ziegler 2007. A new species of the natricine snake genus Amphiesma from the Indochinese Region (Squamata: Colubridae: Natricinae). Zootaxa 1462: 41-60 - get paper here
  • David, P.; Vogel, G. & Pauwels, O.S.G. 2005. On the occurrence of Amphiesma bitaeniatum (WALL, 1925) in Vietnam, with preliminary remarks on the group of Amphiesma parallelum (BOULENGER, 1890) (Serpentes, Colubridae, Natriciniae). Salamandra 41 (4): 167-178 - get paper here
  • DAVID, PATRICK; ISHAN AGARWAL, RAMANA ATHREYA, ROSAMMA MATHEW,<br />GERNOT VOGEL & VIRAL K. MISTRY 2015. Revalidation of Natrix clerki Wall, 1925, an overlooked species in the genus Amphiesma Duméril, Bibron & Duméril, 1854 (Squamata: Natricidae). Zootaxa 3919 (2): 375–395 - get paper here
  • FEI Dong-Bo; YANG Dao-De; SONG Yu-Cheng; FU Qi; YANG Xiang-Lun 2010. New Records of Tropidophorus hainanus and Amphiesma bitaeniata in Hunan,China. Chinese J. Zool. - get paper here
  • GAO, ZHIWEI; TIANYU QIAN, JIANPING JIANG, DEJIA HOU, XUEJIAN DENG & DAODE YANG. 2022. Species diversity and distribution of amphibians and reptiles in Hunan Province, China. Biodiversity Science 30 (2): 21290, 1–15 - get paper here
  • GUO, PENG; FEI ZHU, QIN LIU, LIANG ZHANG, JIAN X. LI, YU Y. HUANG & R. ALEXANDER PYRON 2014. A taxonomic revision of the Asian keelback snakes, genus Amphiesma (Serpentes: Colubridae: Natricinae), with description of a new species. Zootaxa 3873 (4): 425–440 - get paper here
  • Hikida, Tsutomu 2015. Book Review: A Field Guide to the Reptiles of Thailand. Nat. Hist. Bull. Siam Soc. 61(1): 41–51
  • Le, Dzung Trung; Anh Ngoc Dao, Dat Thang Pham, Thomas Ziegler, and Truong Quang Nguyen 2018. New records and an updated list of snakes from Yen Bai Province, Vietnam. Herpetology Notes 11: 101-108 - get paper here
  • LIU Qin; ZHONG Guang-Hui; HU Jian ; ZHAO Hui; GUO Peng 2011. Amphiesma bitaeniata: A Snake New to Guizhou Province,China. Chinese J. Zool. - get paper here
  • Lottier, Anne & Patrick David. 2021. Deux espèces nouvelles pour la faune des serpents du Laos. Bull. Soc. Herp. Fr. 179: 05-10 - get paper here
  • Mell,R. 1931. List of Chinese snakes. Lingnan Sci. Jour., Canton, 8 [1929]: 199-219.
  • Nguyen, S.V., Ho, C.T. and Nguyen, T.Q. 2009. Herpetofauna of Vietnam. Chimaira, Frankfurt, 768 pp.
  • Pham AV, Ziegler T, Nguyen TQ 2020. New records and an updated checklist of snakes from Son La Province, Vietnam. Biodiversity Data Journal 8: e52779 - get paper here
  • Wall, FRANK 1925. Notes on snakes collected in Burma in 1924. J. Bombay Nat. Hist. Soc. 30 (4): 805-821. - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wang K, Lyu ZT, Wang J, Qi S, Che J 2022. Updated Checklist and Zoogeographic Division of the Reptilian Fauna of Yunnan Province, China. Biodiversity Science 30 (4): 21326, 1–31 - get paper here
  • Wang, Kai; Jinlong Ren, Hongman Chen, Zhitong Lyu, Xianguang Guo Ke Jiang, Jinmin Chen, Jiatang Li, Peng Guo, Yingyong Wang, Jing Che 2020. The updated checklists of amphibians and reptiles of China. Biodiversity Science 28 (2): 189-218 - get paper here
  • Zhao, E.M. 2006. The snakes of China [in Chinese]. Hefei, China, Anhui Sience & Technology Publ. House, Vol. I, 372 pp., Vol. II (color plates), 280 pp.
  • ZHOU, ZHENGYAN; ZHIYONG SUN, SHUO QI, YUYAN LU, ZHITONG LYU, YINGYONG WANG, PIPENG LI, JIANZHANG MA 2019. A new species of the genus Hebius (Squamata: Colubridae: Natricinae) from Hunan Province, China. Zootaxa 4674 (1): 068–082 - get paper here
  • Ziegler, T., and Quyet, L.K. 2006. A new natricine snake of the genus Amphiesma (Squamata: Colubridae: Natricinae) from the central Truong Son, Vietnam. Zootaxa 1225: 39-56 - get paper here
 
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