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Helicops carinicaudus (WIED-NEUWIED, 1825)

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Higher TaxaColubridae (Dipsadinae), Colubroidea, Caenophidia, Alethinophidia, Serpentes, Squamata (snakes) 
SubspeciesHelicops carinicaudus carinicaudus WIED-NEUWIED 1825
Helicops carinicaudus triserialis LINDHOLM in LAMPE & LINDHOLM 1902 
Common NamesE: Wied's Keelback
G: Kielschwanznatter 
SynonymColuber carinicaudus WIED-NEUWIED 1825: 300
Homalopsis carinicaudus — SCHLEGEL 1837: 350
Helicops carinicaudus — WAGLER 1830: 170
Helicops carinicaudus — GÜNTHER 1861: 427
Helicops carinicaudus — DUMÉRIL, BIBRON & DUMÉRIL 1854: 744
Helicops carinicaudus — BOULENGER 1886: 435
Helicops carinicauda — BOULENGER 1893: 276
Helicops carinicaudus — PETERS et al. 1970: 123
Helicops carinicaudus — PONTES et al. 2008
Helicops carinicaudus — WALLACH et al. 2014: 321 
DistributionBrazil (Espirito Santo, Rio Grande do Sul, Rio de Janeiro)

triserialis: Type locality: Brasilien [Brasil];  
Reproductionviviparous (Braz et al. 2016) 
TypesHolotype: AMNH 3365, a 909 mm specimen.
Holotype: MWNH 857 (now lost) [triserialis] 
CommentSubspecies: Helicops carinicaudus infrataeniatus is now considered a valid species (see Helicops infrataeniatus). Photos printed in Uruguay (Achaval & Olmos, 2003, 2007, Carreira et al., 2005) present Helicops H. carinicaudus as H. infrataeniatus (T. de Lema, pers. comm., 3 Feb 2017).

Synonymy: Helicops carinicaudus triserialis could presumably represent a junior
synonym of Helicops infrataeniatus (Jan, 1865) (see Hofstadler Deiques & Zanini Cechin 1990, Cei 1993).

Distribution: Not in Argentina according to  GIRAUDO & SCROCCHI 2002. Not in Uruguay according to CARREIRA et al. (2005). One report from Colombia is questionable (Pérez-Santos & Moreno 1988).

Type species: Coluber carinicaudus WIED-NEUWIED 1825 is the type species of the genus Helicops WAGLER 1828. 
References
  • Bérnils, R.S.; Batista, M.A. & Bertelli, P.W. 2001. Cobras e lagartos do Vale: levantamento das espécies de Squamata (Reptilia, Lepidosauria) da Bacia do Rio Itajai, Santa Catarina, Brasil. Rev. Est. Ambientais (Blumenau) 3 (1): 69-79
  • Boulenger, G.A. 1886. A synopsis of the reptiles and batrachians of the province Rio Grande do Sul, Brazil. Ann. Mag. nat. Hist. (5) 18: 423-445 - get paper here
  • Boulenger, G.A. 1893. Catalogue of the snakes in the British Museum (Nat. Hist.) I. London (Taylor & Francis), 448 pp. - get paper here
  • Braz, Henrique B.; Rodrigo R. Scartozzoni, Selma M. Almeida-Santos 2016. Reproductive modes of the South American water snakes: A study system for the evolution of viviparity in squamate reptiles Zoologischer Anzeiger 263: 33-44; doi:10.1016/j.jcz.2016.04.003 - get paper here
  • Castro, T.M. & Teixeira, R.L. 2007. Straßenverkehr als Bedrohung für die Amphibien und Reptilien des Atlantischen Regenwalds in Südost-Brasilien. Eine Fallstudie. Sauria 29 (2): 35-42 - get paper here
  • Costa, Henrique C.; Diego J. Santana, Fernando Leal, Ricardo Koroiva, and Paulo C.A. Garcia 2016. A New Species of Helicops (Serpentes: Dipsadidae: Hydropsini) from Southeastern Brazil. Herpetologica Jun 2016, Vol. 72, No. 2: 157-166
  • Deiques,C.H. & CECHIN,S.T.Z. 1991. O status de Helicops carinicaudus (WIed 1825) (Serpentes: Colubridae). Acta biol. leopoldensia 12 (2): 313-326
  • Freiberg, M.A. 1982. Snakes of South America. T.F.H. Publications, Neptune City, NJ, 189 pp.
  • Giraudo, A. R., and G. J. Scrocchi 2002. Argentinian Snakes: An Annotated Checklist. Smithsonian Herp. Inf. Serv. (132): 1-53 - get paper here
  • Günther, A. 1861. On the ophidian genus Helicops. Ann. Mag. Nat. Hist. (3) 7: 425-428 - get paper here
  • Hamdan, B.; D.P. Coelho; A.B. D’Angiolella; E.J.R. Dias; R.M. Lira-da-Silva. 2013. The Reptile Collection of the Museu de Zoologia, Universidade Federal da Bahia, Brazil. Check List 9 (2):257-262 - get paper here
  • Hoffman, Marlene; Rogério L. Teixeira & Thiago M. Castro 2007. Bemerkungen über die Kielschwanznatter Helicops carinicaudus (WIED-NEUWIED, 1825)(Squamata Colubridae) in einer landwirtschaftlich genutzten Gegend in Südost Brasilien. Ophidia 1 (2) - get paper here
  • Kawashita-Ribeiro, Ricardo Alexandre; Robson Waldemar Ávila, and Drausio Honorio Morais 2013. A New Snake of the Genus Helicops Wagler, 1830 (Dipsadidae, Xenodontinae) from Brazil. Herpetologica 69 (1): 80-90. - get paper here
  • Köhler, J. & Güsten, R. 2007. Herpetological type specimens in the natural history collections of the museums in Darmstadt and Wiesbaden, Germany. Spixiana 30 (2): 275-288 - get paper here
  • Lampe, E. & Lindholm, W.A. 1902. Catalog der Reptilien- und Amphibian-Sammlung (Schlangen; Frosch-, Schwanz- und Schleichenlurche) des Naturhistorischen Museums zu Wiesbaden, mit Bemerkungen. Jahrbuch des Nassauischen Vereins für Naturkunde, Wiesbaden 55: 1–66 - get paper here
  • Oliveira Lula Salles, R. de & Silva-Soares, T. 2010. Répteis do município de Duque de Caxias, Baixada Fluminense, Rio de Janeiro, Sudeste do Brasil. Biotemas, 23 (2): 135-144
  • Pérez-Santos,C. & Moreno, A.G. 1988. Ofidios de Colombia. Museo reegionale di Scienze Naturali, Torino, Monographie VI, 517 pp.
  • Peters, James A.; Donoso-Barros, Roberto & Orejas-Miranda, Braulio 1970. Catalogue of the Neotropical Squamata: Part I Snakes. Part II Lizards and Amphisbaenians. Bull. US Natl. Mus. 297: 347 pp. - get paper here
  • Pontes, J.A.L.; Figueiredo, J.P., Pontes, RC. and Rocha, C.F.D. 2008. Snakes from the Atlantic Rainforest area of Serra do Mendanha, in Rio de Janeiro state, southeastern Brazil: a first approximation to the taxocenosis composition. Braz. J. Biol. 68(3): 601-609 - get paper here
  • ROCHA, CARLOS FREDERICO D.; HELENA G.BERGALLO, JOSÉ P. POMBAL JR., LENA GEISE, MONIQUE VAN SLUYS, RONALDO FERNANDES, ULISSES CARAMASCHI 2004. FAUNA DE ANFÍBIOS, RÉPTEIS E MAMÍFEROS DO ESTADO DO RIO DE JANEIRO, SUDESTE DO BRASIL Publ. Avul. Mus. Nac., Rio de Janeiro (104): 3-23 - get paper here
  • Schlegel, H. 1837. Essai sur la physionomie des serpens. Partie Descriptive. La Haye (J. Kips, J. HZ. et W. P. van Stockum), 606 S. + xvi - get paper here
  • Silva-Soares, T.; R.B. Ferreira; R.O.L. Salles; C.F.D. Rocha. 2011. Continental, insular and coastal marine reptiles from the municipality of Vitória, state of Espírito Santo, southeastern Brazil. Check List 7 (3): 290-298 - get paper here
  • Vrcibradic, D.; C.F.D. Rocha; M.C. Kiefer; F.H. Hatano; A.F. Fontes; M. Almeida-Gomes; C.C. Siqueira; J.A.L. Pontes; V.N.T. Borges-Junior; L.O. Gil; T. Klaion; E.C.N. Rubião; M. Sluys. 2011. Herpetofauna, Estação Ecológica Estadual do Paraíso, state of Rio de Janeiro, southeastern Brazil. Check List 7 (6): 745-749 - get paper here
  • Wallach, Van; Kenneth L. Williams , Jeff Boundy 2014. Snakes of the World: A Catalogue of Living and Extinct Species. [type catalogue] Taylor and Francis, CRC Press, 1237 pp.
  • Wied-Neuwied, M. 1824. Verzeichniss der Amphibien, welche im zweyten Bande der Naturgeschichte Brasiliens vom Prinz Max von Neuwied werden beschrieben werden. Isis von Oken 14: 661—673 [columns] - get paper here
  • Wied-Neuwied, M. Prinz zu 1825. Beiträge zur Naturgeschichte von Brasilien. Weimar, no. 1, xxii + 612 pp. - get paper here
  • Yuki R N. 1994. Regarding Helicops danieli Amaral, 1937, with a description of the hemipenis (Serpentes, Colubridae, Xenodontinae).[in Portugese]. BOLETIM DO MUSEU PARAENSE EMILIO GOELDI SERIE ZOOLOGIA. 1994. vol. 10, no. 2, pp. 203-209.
 
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