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Hemidactylus adensis SMÍD, MORAVEC, KRATOCHVÍL, NASHER, MAZUCH, GVOZDÍK & CARRANZA, 2015

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Higher TaxaGekkonidae, Gekkota, Sauria, Squamata (lizards: geckos) 
Subspecies 
Common Names 
SynonymHemidactylus adensis SMÍD, MORAVEC, KRATOCHVÍL, NASHER, MAZUCH, GVOZDÍK & CARRANZA 2015
Hemidactylus sp. 6 — MORAVEC et al. 2011
Hemidactylus sp. 6 — SMID et al. 2013 
DistributionYemen

Type locality: Yemen, Lahij governorate, Sheikh Othman, 13.917°N, 44.983°E, 22 m elevation  
Reproduction 
TypesHolotype: NHM-BS N41904, adult male (MorphoBank M329154-M329164), 22. III. 2009, collected by S. M. Busais;
Paratypes. NHM-BS N41905-907, subadult male (NHM-BS N41905, MorphoBank M329165-M329172), juvenile (NHM-BS N41906, MorphoBank M329173- M329180), subadult female (NHM-BS N41907, Morpho- Bank M329181-M329189), same data as holotype; NHM-BS N41902-903, adult females MorphoBank M329131- M329140, M329141-M329153, respectively), Yemen, Lahij governorate, Lahij, 13.055°N, 44.878°E, 136 m elevation, 18. I. 2008, collected by S. M. Busais; NMP6V 74837, adult female (MorphoBank M329190-M329206), Yemen, Wadi Tuban (N of Lahij), 13.13°N, 44.85°E, 200 m elevation, 25. X. 2007, collected by L. Kratochvıl. 
CommentDiagnosis. A small-sized Hemidactylus, member of the H. robustus species group as part of the Arabian radiation of the Hemidactylus arid clade characterized by: (1) small size with maximum SVL 37.9 mm for males and 41.8 mm for females; (2) pointy and relatively long and flat head (HL D 24-27% SVL; HD D 40-42% HL); (3) -7 infralabials and 8-10 supralabials; (4) uppermost nasals divided by intervening row of inserted scales; (5) 14 rows of enlarged mostly flat and indistinctly keeled dorsal tubercles; (6) 6-7 preanal pores in males; (7) 5-6 lamellae under the 1st toe; (8) 9-10 lamellae under the 4th toe; (9) enlarged subcaudals; (10) indistinct tail whorls not disrupting the tail outline when viewed from above; (11) contrasting dark brown stripes on the otherwise greyish body running from nostrils over eye and temporal area up to scapular region where they are connected by two to three transverse bars [from SMID et al. 2015].

Sympatry: Hemidactylus flaviviridis, H. y. yerburii, Stenodactylus yemenensis, Acanthodactylus sp.

Distribution: see map in ŠMÍD et al. 2016: Fig. 3. 
EtymologyThe species epithet adensis is an adjective referring to the area where the species is found, lowlands in the vicinity of the city of Aden. 
References
  • MORAVEC, JIŘÍ; LUKÁŠ KRATOCHVÍL, ZUAIR S. AMR, DAVID JANDZIK, JIŘÍ ŠMÍD, & VÁCLAV GVOŽDÍK 2011. High genetic differentiation within the Hemidactylus turcicus complex (Reptilia: Gekkonidae) in the Levant, with comments on the phylogeny and systematics of the genus. Zootaxa 2894: 21–38 - get paper here
  • Šmíd J, Moravec J, Kratochvíl L, Gvoždík V, Nasher AK, Busais SM, Wilms T, Shobrak MY, Carranza S 2013. Two newly recognized species of Hemidactylus (Squamata, Gekkonidae) from the Arabian Peninsula and Sinai, Egypt. ZooKeys 355: 79–107 - get paper here
  • Šmíd, Jiří et al. 2015. Multilocus phylogeny and taxonomic revision of the Hemidactylus robustus species group (Reptilia, Gekkonidae) with descriptions of three new species from Yemen and Ethiopia. Systematics and Biodiversity, DOI: 10.1080/14772000.2014.996264 - get paper here
  • Šmíd, Jiří; Salvador Carranza, Lukáš Kratochvíl, Václav Gvoždík, Abdul Karim Nasher, Jiří Moravec 2013. Out of Arabia: A Complex Biogeographic History of Multiple Vicariance and Dispersal Events in the Gecko Genus Hemidactylus (Reptilia: Gekkonidae). PLoS ONE 8(5): e64018 - get paper here
  • Šmíd, Jiří; Mohammed Shobrak; Thomas Wilms; Ulrich Joger & Salvador Carranza 2016. Endemic diversification in the mountains: genetic, morphological, and geographical differentiation of the Hemidactylus geckos in southwestern Arabia. Org Divers Evol, DOI 10.1007/s13127-016-0293-3
 
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